View source for Chaenactis

{{Treatment/ID
|accepted_name=Chaenactis
|accepted_authority=de Candolle in A. P. de Candolle and A. L. P. P. de Candolle
|publications={{Treatment/Publication
|title=in A. P. de Candolle and A. L. P. P. de Candolle, Prodr.
|place=5: 659. 1836
|year=1836
}}
|common_names=Pincushion;dustymaidens
|basionyms=
|synonyms=
|hierarchy=Asteraceae;Asteraceae tribe Heliantheae;Asteraceae (tribe Heliantheae) subtribe Chaenactidinae;Chaenactis
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Asteraceae]]</div><div class="higher-taxon"><small>tribe</small>[[Asteraceae tribe Heliantheae]]</div><div class="higher-taxon"><small>subtribe</small>[[Asteraceae (tribe Heliantheae) subtribe Chaenactidinae]]</div><div class="higher-taxon"><small>genus</small>[[Chaenactis]]</div></div>
|etymology=Greek chaino, to gape, and aktis, ray, alluding to enlarged peripheral corollas of type species
|volume=Volume 21
|mention_page=page 5, 182, 364, 365, 366, 395, 401, 402
|treatment_page=page 400
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Annuals,</b> biennials, perennials, or subshrubs, (2–)5–70(–200) cm (taprooted). <b>Stems</b> erect to prostrate, usually branched. <b>Leaves</b> basal and/or cauline (smaller and sparser distally except in <i>C. cusickii</i>); alternate; usually petiolate; blades deltate, elliptic, linear, oblanceolate, or ovate (plane or ± 3-dimensional), (0–)1–4-pinnately (rarely -subpalmately) lobed, ultimate margins entire, faces glabrous or hairy, often stipitate-glandular or gland-dotted. <b>Heads</b> discoid or ± radiant, borne singly or in (terminal) ± cymiform arrays (erect in bud except <i>C. macrantha</i>). <b>Involucres</b> hemispheric to obconic or broadly cylindric, (3–)5–15[–25] mm diam. <b>Phyllaries</b> 5–21+ in 1–2(–3) series (subequal to unequal). <b>Receptacles</b> convex to ± flat, pitted and/or knobby, usually epaleate (paleae 3–10+ in <i>C. carphoclinia</i>). <b>Ray</b> florets 0 (sometimes simulated by enlarged peripheral disc corollas). <b>Disc</b> florets 8–70+, bisexual, fertile (diurnal with anthers exserted except in <i>C. macrantha</i>); corollas white, pinkish, cream, or yellow, tubes shorter than cylindric or funnelform throats, lobes 5, deltate to ± lanceolate (sometimes enlarged, unequal; style-branch appendages blunt, obscure). <b>Cypselae</b> clavate to ± cylindric or compressed, obscurely 8–20-angled, faces scabrous and strigose to densely sericeous (usually eglandular); pappi usually persistent, of (1–)4–20, distinct, ± erose scales in 1–4 series (equal or unequal, outer then shorter, scales usually fewer and/or shorter on peripheral cypselae, midnerves obscure), sometimes 0 or coroniform. <b>x</b> = ? (n = 6, 8, plus polyploids and dysploid numbers).</span><!--

-->{{Treatment/Body
|distribution=w North America;nw Mexico.
|discussion=<p>Species 18 (17 in the flora).</p><!--
--><p><i>Chaenactis</i> species grow in arid to alpine or Mediterranean climates, usually in open, unstable or early seral habitats (loose sand, scree, talus, shrink-swell clay, fire-adapted vegetation, recent disturbances). Some annual species have been grown in gardens in the eastern United States.</p><!--
--><p>Differences in induments are key to distinguishing some <i>Chaenactis</i> species. Unless otherwise noted in descriptions and key leads, assume for any given plant that indument of proximal leaves and adjacent proximal portions of stems (“proximal indument”) is similar; indument of distal leaves is, likewise, similar to indument of stem portions from which they arise; peduncle indument is denser distally than proximally; and distal peduncle indument is similar to proximal phyllary indument. Unless otherwise noted, phyllary traits apply to the outer series only. Pappus scales may be equal, subequal, or unequal; unequal scales may intergrade (here said to be in gradually unequal series) or may form two, more or less uniform, shorter and longer series (here said to be in abruptly unequal series); subequal scales are said to be in subequal series. Excellent illustrations of most <i>Chaenactis</i> species appeared in L. Abrams and R. S. Ferris (1923–1960, vol. 4) and A. Cronquist (1955).</p><!--
--><p><i>Section</i> Acarphaea is distinctive by its farinose indument and base chromosome number of 8, among other traits; it could prove to be a separate, convergent genus. Natural and artificial hybrids have been documented among some members of sect. <i>Chaenactis</i> (see further discussion there). Reports of hybrids among species of the other two sections are few and doubtful. <i>Chaenactis</i> appears to be most closely related to the monotypic <i>Dimeresia</i> and <i>Orochaenactis</i>, which B. G. Baldwin et al. (2002) treated together as a narrowly circumscribed tribe, Chaenactideae.</p>
|tables=
|references={{Treatment/Reference
|id=mooring1965a
|text=Mooring, J. S. 1965. Chromosome studies in Chaenactis and Chamaechaenactis (Compositae, Helenieae). Brittonia 17: 17–25.
}}{{Treatment/Reference
|id=stockwell1940a
|text=Stockwell, P. 1940. A revision of the genus Chaenactis. Contr. Dudley Herb. 3: 89–167.
}}
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Annuals; proximal indument predominantly ± farinose, not arachnoid; largest leaf blades (2–)3–4-pinnately lobed
|[[Chaenactis sect. Acarphaea|Chaenactis sect. Acarphaea]]
|-id=key-0-1
|1
|Annuals, biennials, perennials, or subshrubs; proximal indument predominantly arachnoid, lanuginose, pannose, stipitate-glandular, or glabrescent, not farinose; largest leaf blades (0–)1–2-pinnately or -subpalmately lobed
|[[#key-0-2| > 2]]
|-id=key-0-2
|2
|Biennials, perennials, or subshrubs (rarely flowering first year); pappi of (8–)10–20 scales in 2–4 equal or gradually unequal series; leaf blades gland-dotted beneath indument
|[[Chaenactis sect. Macrocarphus|Chaenactis sect. Macrocarphus]]
|-id=key-0-2
|2
|Annuals; pappi usually of (1–)4–8(–14) scales in 1, 2 abruptly unequal, or 2–3 gradually unequal series, sometimes 0 or coroniform; leaf blades not gland-dotted (except C. macrantha)
|[[Chaenactis sect. Chaenactis|Chaenactis sect. Chaenactis]]
|}
</div></div><!--

-->{{#Taxon:
name=Chaenactis
|author=James D. Morefield
|authority=de Candolle in A. P. de Candolle and A. L. P. P. de Candolle
|rank=genus
|parent rank=subtribe
|synonyms=
|basionyms=
|family=Asteraceae
|illustrator=Linny Heagy
|distribution=w North America;nw Mexico.
|reference=mooring1965a;stockwell1940a
|publication title=in A. P. de Candolle and A. L. P. P. de Candolle, Prodr.
|publication year=1836
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/f6b125a955440c0872999024f038d74684f65921/coarse_grained_fna_xml/V19-20-21/V21_1009.xml
|tribe=Asteraceae tribe Heliantheae
|subtribe=Asteraceae (tribe Heliantheae) subtribe Chaenactidinae
|genus=Chaenactis
}}<!--

-->[[Category:Treatment]][[Category:Asteraceae (tribe Heliantheae) subtribe Chaenactidinae]]