View source for Rhytidiadelphus

{{Treatment/ID
|accepted_name=Rhytidiadelphus
|accepted_authority=(Limpricht) Warnstorf
|publications={{Treatment/Publication
|title=Krypt.-Fl. Brandenburg
|place=2: 917. 1906
|year=1906
}}
|basionyms={{Treatment/ID/Basionym
|name=Hylocomium subg. Rhytidiadelphus
|authority=Limpricht
|rank=subgenus
|publication_title=Laubm. Deutschl.
|publication_place=3: 590. 1901
}}
|synonyms={{Treatment/ID/Synonym
|name=Hylocomiadelphus
|authority=
|rank=genus
}}
|hierarchy=Hylocomiaceae;Rhytidiadelphus
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Hylocomiaceae]]</div><div class="higher-taxon"><small>genus</small>[[Rhytidiadelphus]]</div></div>
|etymology=Genus Rhytidium and Greek adelphos, brother, alluding to relationship
|volume=Volume 28
|mention_page=page 316, 326, 331, 338, 642, 644, 645
|treatment_page=page 330
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>creeping to ascending or erect, 2–8 mm wide across leafy stem, monopodial, irregularly and remotely branched to subpinnate; paraphyllia absent. <b>Stem</b> leaves erect-spreading to squarrose, squarrose-recurved, or falcate-secund, crowded or not, broadly ovate, ovate-triangular, or ovate-lanceolate, sometimes plicate proximally, sometimes rugose, 2.1–4.9 mm; base not or slightly decurrent; margins serrulate to nearly entire proximally, serrulate distally; apex acuminate; costa double, short or to 2/3 leaf length; alar cells not differentiated, or shorter, wider than basal cells; laminal cells smooth or coarsely prorate. <b>Branch</b> leaves ovate to narrowly lanceolate. <b>Capsule</b> horizontal; operculum conic; exostome teeth cross striolate to reticulate; endostome segments broadly perforate.</span><!--

-->{{Treatment/Body
|distribution=North America;Eurasia;Atlantic Islands;Pacific Islands (New Zealand);Australia (Tasmania);cool temperate and boreal zones.
|discussion=<p>Species 5 (5 in the flora).</p><!--
--><p>In recent years, M. S. Ignatov and E. A. Ignatova (2003+, vol. 2) and R. Ochyra and A. Stebel (2008) have advocated placing <i>Rhytidiadelphus triquetrus</i> in its own monospecific genus. Although it is morphologically the most distinctive of the five species of <i>Rhytidiadelphus</i>, evolutionary relationships among the five are not clear. Some molecular studies have suggested a less than close relationship between <i>R. squarrosus</i> and <i>R. triquetrus</i>, but these studies were not directly focused on Hylocomiaceae. On the other hand, the analysis of morphological and anatomical evidence by L. Hedenäs (2004) in his cladistic study of Hylocomiaceae revealed that <i>R. loreus</i> and <i>R. triquetrus</i> share a more recent common ancestor than either does with the other three species. Likewise, ITS and ISSR molecular markers employed by A. Vanderpoorten et al. (2003) delineated two pairs of closely related species: <i>R. loreus</i> with <i>R. triquetrus</i> and <i>R. squarrosus</i> with <i>R. subpinnatus</i>. Until further study clarifies relationships within both <i>Rhytidiadelphus</i> and Hylocomiaceae, it seems best to maintain the traditional circumscription of the genus.</p>
|tables=
|references={{Treatment/Reference
|id=koponen1971a
|text=Koponen, T. J. 1971. Rhytidiadelphus japonicus and R. subpinnatus. Hikobia 6: 18–35.
}}{{Treatment/Reference
|id=vanderpoorten2003a
|text=Vanderpoorten, A. et al. 2003. Differentiation in DNA fingerprinting and morphology among species of the pleurocarpous moss genus, Rhytidiadelphus (Hylocomiaceae). Taxon 52: 229–236.
}}
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Leaves plicate; alar cells not or slightly differentiated
|[[#key-0-2| > 2]]
|-id=key-0-1
|1
|Leaves not plicate; alar cells differentiated
|[[#key-0-3| > 3]]
|-id=key-0-2
|2
|Costae 1/2 -2/3 leaf length; medial laminal cells coarsely prorate; stem leaf apices gradually narrowed, acumina broad, flat (occasionally rugose).
|[[Rhytidiadelphus triquetrus|Rhytidiadelphus triquetrus]]
|-id=key-0-2
|2
|Costae less than 1/4 leaf length; medial laminal cells smooth; stem leaf apices abruptly narrowed, acumina slender, channeled.
|[[Rhytidiadelphus loreus|Rhytidiadelphus loreus]]
|-id=key-0-3
|3
|Costae 1/3-1/2 leaf length; alar cell walls thick, porose; stem leaf acumina short.
|[[Rhytidiadelphus japonicus|Rhytidiadelphus japonicus]]
|-id=key-0-3
|3
|Costae to 1/3 leaf length; alar cell walls thin, not porose; stem leaf acumina long
|[[#key-0-4| > 4]]
|-id=key-0-4
|4
|Stem leaves not crowded (stem visible between leaves) except at stem apices, erect-spreading to squarrose; bases not or scarcely sheathing; leaves broadly ovate-triangular; stems irregularly branched to pinnate; moist, usually shaded sites.
|[[Rhytidiadelphus subpinnatus|Rhytidiadelphus subpinnatus]]
|-id=key-0-4
|4
|Stem leaves crowded (stem concealed by leaves), strongly squarrose-recurved; bases subsheathing; leaves ovate; stems remotely branched to irregularly pinnate; well-drained, open to partially shaded sites, near coast or in grassy, disturbed habitats.
|[[Rhytidiadelphus squarrosus|Rhytidiadelphus squarrosus]]
|}
</div></div><!--

-->{{#Taxon:
name=Rhytidiadelphus
|authority=(Limpricht) Warnstorf
|rank=genus
|parent rank=family
|synonyms=Hylocomiadelphus
|basionyms=Hylocomium subg. Rhytidiadelphus
|family=Hylocomiaceae
|distribution=North America;Eurasia;Atlantic Islands;Pacific Islands (New Zealand);Australia (Tasmania);cool temperate and boreal zones.
|reference=koponen1971a;vanderpoorten2003a
|publication title=Krypt.-Fl. Brandenburg
|publication year=1906
|special status=
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V28/V28_514.xml
|genus=Rhytidiadelphus
}}<!--

-->[[Category:Treatment]][[Category:Hylocomiaceae]]