View source for Woodsia obtusa

{{Treatment/ID
|accepted_name=Woodsia obtusa
|accepted_authority=(Swartz) Torrey
|publications={{Treatment/Publication
|title=New York State, Rep. Geol. Surv.
|place=195. 1840
|year=1840
}}
|common_names=Blunt-lobed cliff fern;woodsie à lobes arrondis
|special_status={{Treatment/ID/Special_status
|code=E
|label=Endemic
}}
|basionyms={{Treatment/ID/Basionym
|name=Aspidium obtusum
|authority=Swartz
|rank=species
|publication_title=Syn. Fil.
|publication_place=420. 1806; based on Polypodium obtusum Sprengel, Anleit. Kenntn. Gew. 3: 93. 1804, not Poiret 1804
}}
|synonyms={{Treatment/ID/Synonym
|name=Woodsia perriniana
|authority=(Sprengel) Hooker & Greville
|rank=species
}}
|hierarchy=Dryopteridaceae;Woodsia;Woodsia obtusa
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Dryopteridaceae]]</div><div class="higher-taxon"><small>genus</small>[[Woodsia]]</div><div class="higher-taxon"><small>species</small>[[Woodsia obtusa]]</div></div>
|volume=Volume 2
|mention_page=
|treatment_page=
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact to creeping, erect to horizontal, with few to many persistent petiole bases of unequal lengths; scales often uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. <b>Leaves</b> 8–60 × 2.5–12 cm. <b>Petiole</b> light brown or straw-colored when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered. <b>Blade</b> lanceolate to ovate, 2-pinnate to 2-pinnate-pinnatifid proximally, moderately glandular, rarely somewhat viscid; many glandular hairs with thick stalks and distinctly bulbous tips; rachis with glandular hairs and scattered, often hairlike scales. <b>Pinnae</b> ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex, occasionally attenuate; largest pinnae with 5–14 pairs of pinnules; abaxial and adaxial surfaces glandular, lacking nonglandular hairs or scales. <b>Pinnules</b> dentate, sometimes deeply lobed; margins nonlustrous, thin, with occasional glands, lacking cilia or translucent projections. <b>Vein</b> tips usually enlarged to form whitish hydathodes visible adaxially. <b>Indusia</b> of relatively broad, nonfilamentous segments, these multiseriate throughout, composed of ± isodiametric cells, entire or glandular along distal edge, concealed by or slightly surpassing mature sporangia. <b>Spores</b> averaging 35–47 µm.</span><!--

-->{{Treatment/Body
|distribution=Ont.;Que.;Ala.;Ark.;Conn.;Del.;Fla.;Ga.;Ill.;Ind.;Iowa;Kans.;Ky.;La.;Maine;Mass.;Md.;Mich.;Minn.;Miss.;Mo.;N.C.;N.H.;N.J.;N.Y.;Nebr.;Ohio;Okla.;Pa.;R.I.;S.C.;Tenn.;Tex.;Va.;Vt.;W.Va.;Wis.;only in the flora.
|discussion=<p><i>Woodsia obtusa</i> comprises two cytotypes that are treated here as subspecies because they show subtle morphologic and ecological distinctions and tend to have different distributions. Tetraploid populations (<i></i>subsp.<i> obtusa</i>) are found throughout the eastern flora, commonly occurring on limestone. The diploid (<i></i>subsp.<i> occidentalis</i>) is found near the western edge of the species range, usually on sandstone and granitic substrates. Isozyme studies suggest that <i></i>subsp.<i> obtusa</i> may have been derived from <i></i>subsp.<i> occidentalis</i> through autopolyploidy (M. D. Windham 1993). The westernmost collections of <i>Woodsia obtusa</i> (all <i></i>subsp.<i> occidentalis</i>) come from the Wichita Mountains of Oklahoma and the Edwards Plateau of Texas. Reports of this species from the trans-Pecos region of western Texas are apparently based on misidentifications.</p><!--
--><p>Subspecies 2.</p>
|tables=
|references=
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Spores averaging 42-47 µm; proximal pinnules of lower pinnae usually shallowly lobed or merely dentate; blades coarsely cut and evidently 2-pinnate; stems compact to short-creeping, individual branches usually 5-10 mm diam.
|[[Woodsia obtusa subsp. obtusa|Woodsia obtusa subsp. obtusa]]
|-id=key-0-1
|1
|Spores averaging 35-42 µm; proximal pinnules of lower pinnae usually deeply lobed or pinnatifid; blades finely cut, 2-pinnate-pinnatifid; stems short- to long-creeping, individual branches 3-5 mm diam.
|[[Woodsia obtusa subsp. occidentalis|Woodsia obtusa subsp. occidentalis]]
|}
</div></div><!--

-->{{#Taxon:
name=Woodsia obtusa
|authority=(Swartz) Torrey
|rank=species
|parent rank=genus
|synonyms=Woodsia perriniana
|basionyms=Aspidium obtusum
|family=Dryopteridaceae
|distribution=Ont.;Que.;Ala.;Ark.;Conn.;Del.;Fla.;Ga.;Ill.;Ind.;Iowa;Kans.;Ky.;La.;Maine;Mass.;Md.;Mich.;Minn.;Miss.;Mo.;N.C.;N.H.;N.J.;N.Y.;Nebr.;Ohio;Okla.;Pa.;R.I.;S.C.;Tenn.;Tex.;Va.;Vt.;W.Va.;Wis.;only in the flora.
|reference=None
|publication title=New York State, Rep. Geol. Surv.
|publication year=1840
|special status=Endemic
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_721.xml
|genus=Woodsia
|species=Woodsia obtusa
}}<!--

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[[Category:Treatment]]
[[Category:Woodsia]]
[[Category:Revised Since Print]]