View source for Woodsia oregana

{{Treatment/ID
|accepted_name=Woodsia oregana
|accepted_authority=D. C. Eaton
|publications={{Treatment/Publication
|title=Canad. Naturalist & Quart. J. Sci.
|place=n. s. 2: 90. 1865
|year=1865
}}
|common_names=Oregon cliff fern;woodsie de l'oregon
|special_status={{Treatment/ID/Special_status
|code=F
|label=Illustrated
}}{{Treatment/ID/Special_status
|code=E
|label=Endemic
}}
|basionyms=
|synonyms=
|hierarchy=Dryopteridaceae;Woodsia;Woodsia oregana
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Dryopteridaceae]]</div><div class="higher-taxon"><small>genus</small>[[Woodsia]]</div><div class="higher-taxon"><small>species</small>[[Woodsia oregana]]</div></div>
|volume=Volume 2
|mention_page=
|treatment_page=
}}<!--

--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact, erect to ascending, with few to many persistent petiole bases of unequal lengths; scales often uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. <b>Leaves</b> 4–25 × 1–4 cm. <b>Petiole</b> reddish brown to dark purple proximally when mature, not articulate above base, somewhat pliable and resistant to shattering. <b>Blade</b> linear-lanceolate to narrowly ovate, pinnate-pinnatifid or 2-pinnate proximally, sparsely to moderately glandular, never viscid; glandular hairs with thin stalks and slightly expanded tips; rachis with scattered glandular hairs and occasional hairlike scales. <b>Pinnae</b> ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex; largest pinnae with 3–9 pairs of pinnules; abaxial and adaxial surfaces glabrescent to moderately glandular, lacking nonglandular hairs or scales. <b>Pinnules</b> dentate, often shallowly lobed; margins nonlustrous, thin, with occasional glands, lacking cilia, rarely with 1–2-celled translucent projections. <b>Vein</b> tips slightly (if at all) enlarged, barely visible adaxially. <b>Indusia</b> of narrow, usually filamentous segments, these uniseriate for most of length, composed of ± isodiametric cells, concealed by or slightly surpassing mature sporangia. <b>Spores</b> averaging 39–50 µm.</span><!--

-->{{Treatment/Body
|distribution=Alta.;B.C.;Man.;Ont.;Que.;Sask.;Ariz.;Calif.;Colo.;Idaho;Iowa;Kans.;Mich.;Minn.;Mont.;N.Dak.;N.Mex.;N.Y.;Nebr.;Nev.;Okla.;Oreg.;S.Dak.;Utah;Wash.;Wis.;Wyo.;only in the flora.
|discussion=<p>The variability and promiscuity of <i>Woodsia oregana</i> have been major sources of taxonomic difficulties in <i>Woodsia</i>, and more work will be necessary before relationships in this complex are fully resolved. As defined here, <i>W. oregana</i> comprises two subspecies that are chromosomally and biochemically distinct. In addition, the two taxa are nearly allopatric, with the diploid (<i></i>subsp.<i> oregana</i>) confined to the Pacific Northwest and the tetraploid (<i></i>subsp.<i> cathcartiana</i>) extending from the southwestern United States to eastern Canada. Isozyme studies indicate that <i></i>subsp.<i> cathcartiana</i> is not an autotetraploid derived from known diploid populations of <i></i>subsp.<i> oregana</i>, as was hypothesized by D. F. M. Brown (1964), and it may be more appropriate to recognize these taxa as distinct species. The morphologic features that distinguish these subspecies are very subtle, however, and they are associated primarily with differences in chromosome number. Until further systematic analyses are undertaken, these cytotypes should be maintained as subspecies of <i>W. oregana</i>.</p><!--
--><p>Subspecies 2.</p>
|tables=
|references=
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Spores averaging 39-45 µm; cells on pinnule margins regular in shape, margins appearing entire; adaxial epidermal cells averaging less than 120 µm.
|[[Woodsia oregana subsp. oregana|Woodsia oregana subsp. oregana]]
|-id=key-0-1
|1
|Spores averaging 45-50 µm; cells on pinnule margins irregular in shape, margins usually minutely dentate and appearing ragged; adaxial epidermal cells averaging more than 120 µm.
|[[Woodsia oregana subsp. cathcartiana|Woodsia oregana subsp. cathcartiana]]
|}
</div></div><!--

-->{{#Taxon:
name=Woodsia oregana
|authority=D. C. Eaton
|rank=species
|parent rank=genus
|synonyms=
|basionyms=
|family=Dryopteridaceae
|distribution=Alta.;B.C.;Man.;Ont.;Que.;Sask.;Ariz.;Calif.;Colo.;Idaho;Iowa;Kans.;Mich.;Minn.;Mont.;N.Dak.;N.Mex.;N.Y.;Nebr.;Nev.;Okla.;Oreg.;S.Dak.;Utah;Wash.;Wis.;Wyo.;only in the flora.
|reference=None
|publication title=Canad. Naturalist & Quart. J. Sci.
|publication year=1865
|special status=Illustrated;Endemic
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_24.xml
|genus=Woodsia
|species=Woodsia oregana
}}<!--

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[[Category:Treatment]]
[[Category:Woodsia]]
[[Category:Revised Since Print]]