Difference between revisions of "Cylindropuntia echinocarpa"

(Engelmann & J. M. Bigelow) F. M. Knuth

in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. 1935.

Common names: Silver cholla golden cholla
Basionym: Opuntia echinocarpa Engelmann & J. M. Bigelow Proc. Amer. Acad. Arts. 3: 305. 1856
Synonyms: Cactus echinocarpus (Engelmann & Bigelow) Lemaire Cylindropuntia wigginsii (L. D. Benson) H. Robinson Opuntia wigginsii L. D. Benson
Treatment appears in FNA Volume 4. Treatment on page 115. Mentioned on page 104, 110, 111, 116.
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{{Treatment/ID
 
{{Treatment/ID
 
|accepted_name=Cylindropuntia echinocarpa
 
|accepted_name=Cylindropuntia echinocarpa
|accepted_authority=(Engelmann & J. M. Bigelow) F. M. Knuth in C. Backeberg and F. M. Knuth
+
|accepted_authority=(Engelmann & J. M. Bigelow) F. M. Knuth
 
|publications={{Treatment/Publication
 
|publications={{Treatment/Publication
 
|title=in C. Backeberg and F. M. Knuth, Kaktus-ABC,
 
|title=in C. Backeberg and F. M. Knuth, Kaktus-ABC,
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}}
 
}}
 
|common_names=Silver cholla;golden cholla
 
|common_names=Silver cholla;golden cholla
|basionyms={{Treatment/ID/Synonym
+
|basionyms={{Treatment/ID/Basionym
 
|name=Opuntia echinocarpa
 
|name=Opuntia echinocarpa
 
|authority=Engelmann & J. M. Bigelow
 
|authority=Engelmann & J. M. Bigelow
 +
|rank=species
 +
|publication_title=Proc. Amer. Acad. Arts.
 +
|publication_place=3: 305. 1856
 
}}
 
}}
 
|synonyms={{Treatment/ID/Synonym
 
|synonyms={{Treatment/ID/Synonym
 
|name=Cactus echinocarpus
 
|name=Cactus echinocarpus
 
|authority=(Engelmann & Bigelow) Lemaire
 
|authority=(Engelmann & Bigelow) Lemaire
}}{{Treatment/ID/Synonym
+
|rank=species
 +
}} {{Treatment/ID/Synonym
 
|name=Cylindropuntia wigginsii
 
|name=Cylindropuntia wigginsii
 
|authority=(L. D. Benson) H. Robinson
 
|authority=(L. D. Benson) H. Robinson
}}{{Treatment/ID/Synonym
+
|rank=species
 +
}} {{Treatment/ID/Synonym
 
|name=Opuntia wigginsii
 
|name=Opuntia wigginsii
 
|authority=L. D. Benson
 
|authority=L. D. Benson
 +
|rank=species
 
}}
 
}}
 
|hierarchy=Cactaceae;Cactaceae subfam. Opuntioideae;Cylindropuntia;Cylindropuntia echinocarpa
 
|hierarchy=Cactaceae;Cactaceae subfam. Opuntioideae;Cylindropuntia;Cylindropuntia echinocarpa
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}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="tree orientation;tree architecture;tree some measurement;shrub orientation;shrub architecture;shrub some measurement"><b>Trees </b>or shrubs, spreading, densely branched, 0.5–2 m.</span> <span class="statement" id="st-d0_s1" data-properties="stem segment fixation;terminal segment fusion;terminal segment coloration;terminal segment coloration;terminal segment coloration;terminal segment shape;terminal segment atypical length;terminal segment length;terminal segment width"><b>Stem </b>segments usually firmly attached, terminal segments sometimes easily detached, green to light gray-green, cylindric, 3–7.5 (–12) × 1–2.5 cm;</span> <span class="statement" id="st-d0_s2" data-properties="tubercle orientation;tubercle height or length or size;tubercle shape;tubercle atypical some measurement;tubercle some measurement;tubercle width">tubercles salient, short oval, 0.4–1.3 (–1.5) cm, moderately broad;</span> <span class="statement" id="st-d0_s3" data-properties="areole arrangement or shape;areole length;areole width">areoles broadly elliptic, 4–4.5 × 2–3.5 mm;</span> <span class="statement" id="st-d0_s4" data-properties="wool coloration;wool coloration;wool coloration;wool coloration;wool life cycle;wool coloration">wool whitish, yellow to tan, aging gray.</span> <span class="statement" id="st-d0_s5" data-properties="spine atypical count;spine atypical count;spine count;spine count;spine height or length or size;spine architecture;0-areole prominence;0-areole architecture or shape;areole arrangement"><b>Spines </b>(6–) 10–17 (–22) per areole, accompanied by 0–5 short bristlelike spines at areole margins, at most areoles, most prominent in adaxial ones, much interlaced with those of adjacent areoles, obscuring stems, sometimes brushlike;</span> <span class="statement" id="st-d0_s6" data-properties="abaxial spine size;abaxial spine coloration;abaxial spine coloration;abaxial spine coloration;abaxial spine coloration;abaxial spine shape;abaxial spine shape;abaxial spine shape;abaxial spine orientation;abaxial spine orientation;abaxial spine orientation;abaxial spine length;abaxial spine atypical some measurement;abaxial spine some measurement">major abaxial spines whitish, yellow to tan, usually subterete to flattened, spreading to deflexed, the longest 20–30 (–45) mm;</span> <span class="statement" id="st-d0_s7" data-properties="adaxial spine size;adaxial spine orientation;adaxial spine orientation;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine shape;adaxial spine length;adaxial spine atypical some measurement;adaxial spine some measurement">major adaxial spines erect or spreading, white, pale-yellow, or tan to brown (rarely deep redbrown), terete, the longest 22–35 (–50) mm;</span> <span class="statement" id="st-d0_s8" data-properties="sheath coloration;tip coloration;tip coloration;tip coloration">sheaths whitish with yellow to golden tips, baggy.</span> <span class="statement" id="st-d0_s9" data-properties="glochid prominence"><b>Glochids </b>conspicuous in broad adaxial crescent, yellow, 3–4 (–5) mm.</span> <span class="statement" id="st-d0_s10" data-properties="inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal shape;inner tepal some measurement;inner tepal architecture or shape"><b>Flowers:</b> inner tepals light green to yellow-green, sometimes suffused with maroon or rose, spatulate, 20–23 mm, emarginate-apiculate;</span> <span class="statement" id="st-d0_s11" data-properties="filament coloration;filament coloration;filament coloration;filament coloration;filament coloration">filaments greenish white or yellow, sometimes suffused with bronze or rose, but not red;</span> <span class="statement" id="st-d0_s12" data-properties="anther coloration">anthers yellow;</span> <span class="statement" id="st-d0_s13" data-properties="style coloration;style coloration;style coloration;style coloration;stigma lobe coloration;stigma lobe coloration;stigma lobe coloration;stigma lobe coloration">style and stigma lobes whitish, cream to light green.</span> <span class="statement" id="st-d0_s14" data-properties="fruit coloration;fruit shape;fruit shape;fruit shape;fruit length;fruit width;fruit condition or texture;fruit relief;fruit architecture or shape;fruit architecture or shape;umbilicus depth"><b>Fruits </b>tan at maturity, obconic to subspheric, 13–23 × 11–20 mm, dry, tuberculate, densely spiny, sometimes burlike, with apical flange around deep umbilicus;</span> <span class="statement" id="st-d0_s15" data-properties="basal tubercle length or size">basal tubercles not markedly longer than distal ones;</span> <span class="statement" id="st-d0_s16" data-properties="areole atypical count;areole count">areoles 16–24 (–36).</span> <span class="statement" id="st-d0_s17" data-properties="seed coloration;seed shape;seed shape;seed shape;seed shape;seed length;seed width;side architecture or pubescence or relief;depression count;depression size"><b>Seeds </b>pale-yellow, angular to squarish in outline, warped, 4–6 × 3.5–5 mm, sides smooth, each with 2–4 large depressions;</span> <span class="statement" id="st-d0_s18" data-properties="">girdle smooth (rarely with very narrow ridge).</span> <span class="statement" id="st-d0_s19" data-properties="girdle architecture or pubescence or relief;2n chromosome count">2n = 22.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Trees </b>or shrubs, spreading, densely branched, 0.5–2 m. <b>Stem</b> segments usually firmly attached, terminal segments sometimes easily detached, green to light gray-green, cylindric, 3–7.5(–12) × 1–2.5 cm; tubercles salient, short oval, 0.4–1.3(–1.5) cm, moderately broad; areoles broadly elliptic, 4–4.5 × 2–3.5 mm; wool whitish, yellow to tan, aging gray. <b>Spines</b> (6–)10–17(–22) per areole, accompanied by 0–5 short bristlelike spines at areole margins, at most areoles, most prominent in adaxial ones, much interlaced with those of adjacent areoles, obscuring stems, sometimes brushlike; major abaxial spines whitish, yellow to tan, usually subterete to flattened, spreading to deflexed, the longest 20–30(–45) mm; major adaxial spines erect or spreading, white, pale yellow, or tan to brown (rarely deep red-brown), terete, the longest 22–35(–50) mm; sheaths whitish with yellow to golden tips, baggy. <b>Glochids</b> conspicuous in broad adaxial crescent, yellow, 3–4(–5) mm. <b>Flowers</b>: inner tepals light green to yellow-green, sometimes suffused with maroon or rose, spatulate, 20–23 mm, emarginate-apiculate; filaments greenish white or yellow, sometimes suffused with bronze or rose, but not red; anthers yellow; style and stigma lobes whitish, cream to light green. <b>Fruits</b> tan at maturity, obconic to subspheric, 13–23 × 11–20 mm, dry, tuberculate, densely spiny, sometimes burlike, with apical flange around deep umbilicus; basal tubercles not markedly longer than distal ones; areoles 16–24(–36). <b>Seeds</b> pale yellow, angular to squarish in outline, warped, 4–6 × 3.5–5 mm, sides smooth, each with 2–4 large depressions; girdle smooth (rarely with very narrow ridge). <b>2n</b> = 22.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
 +
|phenology=Flowering spring (Mar–Jun).
 
|habitat=Mojave and Sonoran deserts, desert grasslands, juniper and oak-juniper woodlands, flats, bajadas, canyons, sandy, loam, alluvial to gravelly substrates
 
|habitat=Mojave and Sonoran deserts, desert grasslands, juniper and oak-juniper woodlands, flats, bajadas, canyons, sandy, loam, alluvial to gravelly substrates
 
|elevation=50-1700 m
 
|elevation=50-1700 m
 
|distribution=Ariz.;Calif.;Nev.;Mexico (Baja California;Sonora).
 
|distribution=Ariz.;Calif.;Nev.;Mexico (Baja California;Sonora).
|discussion=<p>A dwarf form, with narrow terminal stem segments bearing few spines per areole, and one spine longer than the others, occurs on the very arid flats along the lower Colorado River in California and Arizona and has been recognized as Opuntia wigginsii L. D. Benson; however, numerous intermediate to more robust forms make recognition of the dwarf form untenable. Although exceptions occur, plants to the north and west in the range tend to be more spiny and to bear yellow to yellow-green flowers.</p><!--
+
|discussion=<p>A dwarf form, with narrow terminal stem segments bearing few spines per areole, and one spine longer than the others, occurs on the very arid flats along the lower Colorado River in California and Arizona and has been recognized as <i>Opuntia</i> wigginsii L. D. Benson; however, numerous intermediate to more robust forms make recognition of the dwarf form untenable. Although exceptions occur, plants to the north and west in the range tend to be more spiny and to bear yellow to yellow-green flowers.</p><!--
--><p>Cylindropuntia echinocarpa hybridizes with C. acanthocarpa [= C. ×deserta (Griffiths) Pinkava (D. J. Pinkava 1999)], C. whipplei, and C. munzii. Cylindropuntia ×deserta, found in western Arizona and southern California, has reddish filaments, short tubercles, variably sized spines/areole, and dry spiny fruits.</p>
+
--><p><i>Cylindropuntia echinocarpa</i> hybridizes with <i>C. acanthocarpa</i> [= C. ×deserta (Griffiths) Pinkava (D. J. Pinkava 1999)], <i>C. whipplei</i>, and <i>C. munzii</i>. <i>Cylindropuntia</i> ×deserta, found in western Arizona and southern California, has reddish filaments, short tubercles, variably sized spines/areole, and dry spiny fruits.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Cylindropuntia echinocarpa
 
name=Cylindropuntia echinocarpa
|author=
+
|authority=(Engelmann & J. M. Bigelow) F. M. Knuth
|authority=(Engelmann & J. M. Bigelow) F. M. Knuth in C. Backeberg and F. M. Knuth
 
 
|rank=species
 
|rank=species
 
|parent rank=genus
 
|parent rank=genus
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|basionyms=Opuntia echinocarpa
 
|basionyms=Opuntia echinocarpa
 
|family=Cactaceae
 
|family=Cactaceae
 +
|phenology=Flowering spring (Mar–Jun).
 
|habitat=Mojave and Sonoran deserts, desert grasslands, juniper and oak-juniper woodlands, flats, bajadas, canyons, sandy, loam, alluvial to gravelly substrates
 
|habitat=Mojave and Sonoran deserts, desert grasslands, juniper and oak-juniper woodlands, flats, bajadas, canyons, sandy, loam, alluvial to gravelly substrates
 
|elevation=50-1700 m
 
|elevation=50-1700 m
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|publication year=1935
 
|publication year=1935
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_221.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V4/V4_221.xml
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|genus=Cylindropuntia
 
|genus=Cylindropuntia
 
|species=Cylindropuntia echinocarpa
 
|species=Cylindropuntia echinocarpa
|0-areole architecture or shape=brushlike
 
|0-areole prominence=prominent
 
|2n chromosome count=22
 
|abaxial spine atypical some measurement=30mm;45mm
 
|abaxial spine coloration=yellow;tan
 
|abaxial spine length=longest
 
|abaxial spine orientation=spreading;deflexed
 
|abaxial spine shape=usually subterete;flattened
 
|abaxial spine size=major
 
|abaxial spine some measurement=20mm;30mm
 
|adaxial spine atypical some measurement=35mm;50mm
 
|adaxial spine coloration=tan;brown
 
|adaxial spine length=longest
 
|adaxial spine orientation=spreading;erect
 
|adaxial spine shape=terete
 
|adaxial spine size=major
 
|adaxial spine some measurement=22mm;35mm
 
|anther coloration=yellow
 
|areole arrangement=adjacent
 
|areole arrangement or shape=elliptic
 
|areole atypical count=24;36
 
|areole count=16;24
 
|areole length=4mm;4.5mm
 
|areole width=2mm;3.5mm
 
|basal tubercle length or size=not markedly longer
 
|depression count=2;4
 
|depression size=large
 
|filament coloration=not red;,;suffused with bronze or suffused with rose;yellow;greenish white
 
|fruit architecture or shape=burlike;spiny
 
|fruit coloration=tan
 
|fruit condition or texture=dry
 
|fruit length=13mm;23mm
 
|fruit relief=tuberculate
 
|fruit shape=obconic;subspheric
 
|fruit width=11mm;20mm
 
|girdle architecture or pubescence or relief=smooth
 
|glochid prominence=conspicuous
 
|inner tepal architecture or shape=emarginate-apiculate
 
|inner tepal coloration=,;suffused with maroon or suffused with rose;light green;yellow-green
 
|inner tepal shape=spatulate
 
|inner tepal some measurement=20mm;23mm
 
|seed coloration=pale-yellow
 
|seed length=4mm;6mm
 
|seed shape=warped;angular;squarish
 
|seed width=3.5mm;5mm
 
|sheath coloration=whitish
 
|shrub architecture=branched
 
|shrub orientation=spreading
 
|shrub some measurement=0.5m;2m
 
|side architecture or pubescence or relief=smooth
 
|spine architecture=bristlelike
 
|spine atypical count=17;22
 
|spine count=0;5
 
|spine height or length or size=short
 
|stem segment fixation=attached
 
|stigma lobe coloration=cream;light green
 
|style coloration=cream;light green
 
|terminal segment atypical length=7.5cm;12cm
 
|terminal segment coloration=green;light gray-green
 
|terminal segment fusion=detached
 
|terminal segment length=3cm;7.5cm
 
|terminal segment shape=cylindric
 
|terminal segment width=1cm;2.5cm
 
|tip coloration=yellow;golden
 
|tree architecture=branched
 
|tree orientation=spreading
 
|tree some measurement=0.5m;2m
 
|tubercle atypical some measurement=1.3cm;1.5cm
 
|tubercle height or length or size=short
 
|tubercle orientation=salient
 
|tubercle shape=oval
 
|tubercle some measurement=0.4cm;1.3cm
 
|tubercle width=broad
 
|umbilicus depth=deep
 
|wool coloration=gray;yellow;tan
 
|wool life cycle=aging
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Cylindropuntia]]
 
-->[[Category:Treatment]][[Category:Cylindropuntia]]

Latest revision as of 22:57, 5 November 2020

Trees or shrubs, spreading, densely branched, 0.5–2 m. Stem segments usually firmly attached, terminal segments sometimes easily detached, green to light gray-green, cylindric, 3–7.5(–12) × 1–2.5 cm; tubercles salient, short oval, 0.4–1.3(–1.5) cm, moderately broad; areoles broadly elliptic, 4–4.5 × 2–3.5 mm; wool whitish, yellow to tan, aging gray. Spines (6–)10–17(–22) per areole, accompanied by 0–5 short bristlelike spines at areole margins, at most areoles, most prominent in adaxial ones, much interlaced with those of adjacent areoles, obscuring stems, sometimes brushlike; major abaxial spines whitish, yellow to tan, usually subterete to flattened, spreading to deflexed, the longest 20–30(–45) mm; major adaxial spines erect or spreading, white, pale yellow, or tan to brown (rarely deep red-brown), terete, the longest 22–35(–50) mm; sheaths whitish with yellow to golden tips, baggy. Glochids conspicuous in broad adaxial crescent, yellow, 3–4(–5) mm. Flowers: inner tepals light green to yellow-green, sometimes suffused with maroon or rose, spatulate, 20–23 mm, emarginate-apiculate; filaments greenish white or yellow, sometimes suffused with bronze or rose, but not red; anthers yellow; style and stigma lobes whitish, cream to light green. Fruits tan at maturity, obconic to subspheric, 13–23 × 11–20 mm, dry, tuberculate, densely spiny, sometimes burlike, with apical flange around deep umbilicus; basal tubercles not markedly longer than distal ones; areoles 16–24(–36). Seeds pale yellow, angular to squarish in outline, warped, 4–6 × 3.5–5 mm, sides smooth, each with 2–4 large depressions; girdle smooth (rarely with very narrow ridge). 2n = 22.


Phenology: Flowering spring (Mar–Jun).
Habitat: Mojave and Sonoran deserts, desert grasslands, juniper and oak-juniper woodlands, flats, bajadas, canyons, sandy, loam, alluvial to gravelly substrates
Elevation: 50-1700 m

Distribution

V4 221-distribution-map.gif

Ariz., Calif., Nev., Mexico (Baja California, Sonora).

Discussion

A dwarf form, with narrow terminal stem segments bearing few spines per areole, and one spine longer than the others, occurs on the very arid flats along the lower Colorado River in California and Arizona and has been recognized as Opuntia wigginsii L. D. Benson; however, numerous intermediate to more robust forms make recognition of the dwarf form untenable. Although exceptions occur, plants to the north and west in the range tend to be more spiny and to bear yellow to yellow-green flowers.

Cylindropuntia echinocarpa hybridizes with C. acanthocarpa [= C. ×deserta (Griffiths) Pinkava (D. J. Pinkava 1999)], C. whipplei, and C. munzii. Cylindropuntia ×deserta, found in western Arizona and southern California, has reddish filaments, short tubercles, variably sized spines/areole, and dry spiny fruits.

Selected References

None.

Lower Taxa

None.
Donald J. Pinkava +
(Engelmann & J. M. Bigelow) F. M. Knuth +
Opuntia echinocarpa +
Silver cholla +  and golden cholla +
Ariz. +, Calif. +, Nev. +, Mexico (Baja California +  and Sonora). +
50-1700 m +
Mojave and Sonoran deserts, desert grasslands, juniper and oak-juniper woodlands, flats, bajadas, canyons, sandy, loam, alluvial to gravelly substrates +
Flowering spring (Mar–Jun). +
in C. Backeberg and F. M. Knuth, Kaktus-ABC, +
Cactus echinocarpus +, Cylindropuntia wigginsii +  and Opuntia wigginsii +
Cylindropuntia echinocarpa +
Cylindropuntia +
species +