Difference between revisions of "Diphasiastrum"

Holub

Preslia 47: 104. 1975.

Common names: False Diphasium
Etymology: Diphasium, a generic name, and -astrum, incomplete resemblance
Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties="plant growth form or habitat;plant orientation;shoot orientation"><b>Plants </b>terrestrial, mainly trailing on ground with erect shoots.</span> <span class="statement" id="st-d0_s1" data-properties="root position relational"><b>Roots </b>emerging immediately on underside of main-stems.</span> <span class="statement" id="st-d0_s2" data-properties="stem orientation;stem growth form or orientation;surface location"><b>Horizontal </b>stems on substrate surface to subterranean, long-creeping.</span> <span class="statement" id="st-d0_s3" data-properties="shoot orientation;shoot shape;shoot shape;shoot shape;shoot diameter;lateral branchlet count;main stem orientation"><b>Upright </b>shoots quadrate to flattened (except in D. sitchense), 2–6 mm diam., usually with 2–5 lateral branchlets on main erect stem;</span> <span class="statement" id="st-d0_s4" data-properties="lateral branchlet architecture;lateral branchlet prominence or shape">lateral branchlets leafy, ± flat in cross-section.</span> <span class="statement" id="st-d0_s5" data-properties="leaf fixation or orientation;leaf shape;leaf shape;leaf shape;leaf width;leaf shape;stem orientation;stem arrangement"><b>Leaves </b>on horizontal stems somewhat distant, appressed, linear to lanceolate, thin, scalelike;</span> <span class="statement" id="st-d0_s6" data-properties="leaf arrangement;leaf shape;leaf shape;leaf shape;leaf shape;leaf arrangement;ultimate branchlet fixation or orientation;rank count;leaf size">leaves on ultimate branchlets appressed to divergent, linear-lanceolate to nearly filiform, usually almost scalelike and mostly imbricate, in 4 ranks, leaves of lateral ranks larger, more spreading than those of upperside and underside ranks (except in Diphasiastrum sitchense with 5 ranks of uniform nonimbricate leaves).</span> <span class="statement" id="st-d0_s7" data-properties="branchlet architecture;branchlet count;gemma architecture;gemma count"><b>Gemmiferous </b>branchlets and gemmae absent.</span> <span class="statement" id="st-d0_s8" data-properties="strobilus architecture or arrangement or growth form;strobilus architecture;strobilus count;strobilus architecture;apex shape;apex shape;apical projection reproduction"><b>Strobili </b>solitary and sessile or multiple and stalked, apex blunt, acute, or with sterile apical projection;</span> <span class="statement" id="st-d0_s9" data-properties="peduncle architecture">peduncle conspicuously leafy;</span> <span class="statement" id="st-d0_s10" data-properties="sporophyll height or length or size">sporophylls shorter than peduncle leaves.</span> <span class="statement" id="st-d0_s11" data-properties="sporangium shape"><b>Sporangia </b>reniform.</span> <span class="statement" id="st-d0_s12" data-properties="spore architecture or coloration or relief;side shape;angle shape"><b>Spores </b>reticulate, sides at equator convex, angles acute.</span> <span class="statement" id="st-d0_s13" data-properties="gametophyte structure in adjective form;gametophyte shape;paraphyse count"><b>Gametophytes </b>nonphotosynthetic, mycorrhizal, carrot-shaped, paraphyses absent;</span> <span class="statement" id="st-d0_s14" data-properties="">ring meristem present.</span> <span class="statement" id="st-d0_s15" data-properties="ring meristem count;x chromosome count">x = 23.</span><!--
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--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>terrestrial, mainly trailing on ground with erect shoots. <b>Roots</b> emerging immediately on underside of main stems. <b>Horizontal</b> stems on substrate surface to subterranean, long-creeping. <b>Upright</b> shoots quadrate to flattened (except in <i>D. sitchense</i>), 2–6 mm diam., usually with 2–5 lateral branchlets on main erect stem; lateral branchlets leafy, ± flat in cross section. <b>Leaves</b> on horizontal stems somewhat distant, appressed, linear to lanceolate, thin, scalelike; leaves on ultimate branchlets appressed to divergent, linear-lanceolate to nearly filiform, usually almost scalelike and mostly imbricate, in 4 ranks, leaves of lateral ranks larger, more spreading than those of upperside and underside ranks (except in <i>Diphasiastrum sitchense</i> with 5 ranks of uniform nonimbricate leaves). <b>Gemmiferous</b> branchlets and gemmae absent. <b>Strobili</b> solitary and sessile or multiple and stalked, apex blunt, acute, or with sterile apical projection; peduncle conspicuously leafy; sporophylls shorter than peduncle leaves. <b>Sporangia</b> reniform. <b>Spores</b> reticulate, sides at equator convex, angles acute. <b>Gametophytes</b> nonphotosynthetic, mycorrhizal, carrot-shaped, paraphyses absent; ring meristem present. <b>x</b> = 23.</span><!--
  
 
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|distribution=Mainly north temperate and subarctic.
 
|distribution=Mainly north temperate and subarctic.
|discussion=<p>Diphasiastrum is remarkable in North America for its ability to form numerous homoploid, apparently fertile interspecific hybrids, some of which are frequent to common and must be reckoned with in floristic studies. Field and laboratory studies of these hybrids and their parents are needed for determination of the population dynamics of Diphasiastrum and to answer questions as to why the parental species retain their genetic identities.</p><!--
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|discussion=<p><i>Diphasiastrum</i> is remarkable in North America for its ability to form numerous homoploid, apparently fertile interspecific hybrids, some of which are frequent to common and must be reckoned with in floristic studies. Field and laboratory studies of these hybrids and their parents are needed for determination of the population dynamics of <i>Diphasiastrum</i> and to answer questions as to why the parental species retain their genetic identities.</p><!--
 
--><p>Species 15–20 (11 taxa in the flora, including 5 species and 6 fertile hybrids).</p>
 
--><p>Species 15–20 (11 taxa in the flora, including 5 species and 6 fertile hybrids).</p>
 
|tables=
 
|tables=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Diphasiastrum
 
name=Diphasiastrum
|author=
 
 
|authority=Holub
 
|authority=Holub
 
|rank=genus
 
|rank=genus
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|publication year=1975
 
|publication year=1975
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_502.xml
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|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_502.xml
 
|genus=Diphasiastrum
 
|genus=Diphasiastrum
|angle shape=acute
 
|apex shape=acute;blunt
 
|apical projection reproduction=sterile
 
|branchlet architecture=gemmiferous
 
|branchlet count=absent
 
|gametophyte shape=carrot-shaped
 
|gametophyte structure in adjective form=mycorrhizal
 
|gemma architecture=gemmiferous
 
|gemma count=absent
 
|lateral branchlet architecture=leafy
 
|lateral branchlet count=2;5
 
|lateral branchlet prominence or shape=flat
 
|leaf arrangement=imbricate;divergent
 
|leaf fixation or orientation=appressed
 
|leaf shape=scalelike;linear-lanceolate;nearly filiform
 
|leaf size=larger
 
|leaf width=thin
 
|main stem orientation=erect
 
|paraphyse count=absent
 
|peduncle architecture=leafy
 
|plant growth form or habitat=terrestrial
 
|plant orientation=trailing
 
|rank count=4
 
|ring meristem count=present
 
|root position relational=emerging
 
|shoot diameter=2mm;6mm
 
|shoot orientation=upright;erect
 
|shoot shape=quadrate;flattened
 
|side shape=convex
 
|sporangium shape=reniform
 
|spore architecture or coloration or relief=reticulate
 
|sporophyll height or length or size=shorter
 
|stem arrangement=distant
 
|stem growth form or orientation=long-creeping
 
|stem orientation=horizontal;horizontal
 
|strobilus architecture=stalked;sessile
 
|strobilus architecture or arrangement or growth form=solitary
 
|strobilus count=multiple
 
|surface location=subterranean
 
|ultimate branchlet fixation or orientation=appressed
 
|x chromosome count=23
 
 
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Latest revision as of 21:23, 5 November 2020

Plants terrestrial, mainly trailing on ground with erect shoots. Roots emerging immediately on underside of main stems. Horizontal stems on substrate surface to subterranean, long-creeping. Upright shoots quadrate to flattened (except in D. sitchense), 2–6 mm diam., usually with 2–5 lateral branchlets on main erect stem; lateral branchlets leafy, ± flat in cross section. Leaves on horizontal stems somewhat distant, appressed, linear to lanceolate, thin, scalelike; leaves on ultimate branchlets appressed to divergent, linear-lanceolate to nearly filiform, usually almost scalelike and mostly imbricate, in 4 ranks, leaves of lateral ranks larger, more spreading than those of upperside and underside ranks (except in Diphasiastrum sitchense with 5 ranks of uniform nonimbricate leaves). Gemmiferous branchlets and gemmae absent. Strobili solitary and sessile or multiple and stalked, apex blunt, acute, or with sterile apical projection; peduncle conspicuously leafy; sporophylls shorter than peduncle leaves. Sporangia reniform. Spores reticulate, sides at equator convex, angles acute. Gametophytes nonphotosynthetic, mycorrhizal, carrot-shaped, paraphyses absent; ring meristem present. x = 23.

Distribution

Mainly north temperate and subarctic.

Discussion

Diphasiastrum is remarkable in North America for its ability to form numerous homoploid, apparently fertile interspecific hybrids, some of which are frequent to common and must be reckoned with in floristic studies. Field and laboratory studies of these hybrids and their parents are needed for determination of the population dynamics of Diphasiastrum and to answer questions as to why the parental species retain their genetic identities.

Species 15–20 (11 taxa in the flora, including 5 species and 6 fertile hybrids).

Key

1 Upright shoots to 18 cm, mostly less than 12 cm including base; strobili mostly sessile. > 2
1 Upright shoots, 8–50 cm, mostly more than 12 cm, including base; strobili stalked. > 3
2 Ultimate branchlets approximately square in cross section; leaves 4-ranked, strongly overlapping. Diphasiastrum alpinum
2 Ultimate branchlets round in cross section; leaves 5-ranked, not overlapping. Diphasiastrum sitchense
3 Ultimate branchlets cordlike, nearly square in cross section, usually bluish glaucous; underside leaves approximately equal in size to lateral and upperside leaves. Diphasiastrum tristachyum
3 Ultimate branchlets narrowly bladelike, flat in cross section, usually green; underside leaves much smaller than lateral and upperside leaves. > 4
4 Branchlets irregular, with conspicuous annual bud constrictions; peduncles, if present, regularly forked; strobili mostly 15–25 mm, lacking sterile tips. Diphasiastrum complanatum
4 Branchlets very regularly fan-shaped, lacking conspicuous annual bud constrictions; peduncles mostly branching abruptly at base to produce false whorl of strobili; strobili mostly 20–35 mm, many with sterile tips. Diphasiastrum digitatum