Difference between revisions of "Echinocereus triglochidiatus"

Engelmann in F. A. Wislizenus

in F. A. Wislizenus, Mem. Tour N. Mexico, 93. 1848.

Common names: Claret-cup cactus kingcup cactus
Synonyms: Echinocereus triglochidiatus var. gonacanthus (Engelmann & J. M. Bigelow) BoissevainEchinocereus triglochidiatus var. inermis (Schumann) ArpEchinocereus triglochidiatus var. mojavensis (Engelmann & J. M. Bigelow) L. D. Benson
Treatment appears in FNA Volume 4. Mentioned on page 168.
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--><span class="statement" id="st-d0_s0" data-properties="plant architecture;plant architecture;mound size;branch count"><b>Plants </b>unbranched or 1–12 (–76) -branched, forming large mounds of branches to 300.</span> <span class="statement" id="st-d0_s1" data-properties="stem orientation;stem orientation;stem shape;stem atypical length;stem length;stem atypical width;stem width"><b>Stems </b>usually erect or nearly so, cylindric (spheric), (2–) 5–70 × (3–) 5–13 cm;</span> <span class="statement" id="st-d0_s2" data-properties="rib count;rib count;crest shape">ribs 5–8 or 8–12, crests slightly undulate (localized populations contain plants with strongly interrupted ribs);</span> <span class="statement" id="st-d0_s3" data-properties="areole some measurement;areole arrangement">areoles 10–40 mm apart.</span> <span class="statement" id="st-d0_s4" data-properties="spine atypical count;spine count;spine course;spine course;spine course;spine arrangement or shape;spine orientation;spine orientation;spine orientation;spine orientation;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration"><b>Spines </b>(0–) 3–11 per areole, straight to curved or contorted, appressed (radial spines) or spreading to projecting outward (some radials and central spines when present), white to yellow, gray, or black;</span> <span class="statement" id="st-d0_s5" data-properties="spine arrangement;spine atypical count;spine count;spine atypical some measurement;spine some measurement">radial spines (0–) 1–10 per areole, (0–) 15–90 mm;</span> <span class="statement" id="st-d0_s6" data-properties="central spine atypical count;central spine count;central spine arrangement or shape;central spine atypical some measurement;central spine some measurement">central spines 0–1 (–4) per areole, angular, (0–) 50–120 mm.</span> <span class="statement" id="st-d0_s7" data-properties="flower atypical length;flower length;flower width"><b>Flowers </b>(4–) 5–10 × 3–7 cm;</span> <span class="statement" id="st-d0_s8" data-properties="flower tube some measurement">flower tube 20–35 mm;</span> <span class="statement" id="st-d0_s9" data-properties="flower tube hair some measurement">flower tube hairs 1–2 mm;</span> <span class="statement" id="st-d0_s10" data-properties="inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal atypical length;inner tepal length;inner tepal atypical width;inner tepal width;tip width;tip texture">inner tepals bright-orange-red to dark red, proximally paler (bases sometimes yellow or white), (18–) 25–40 × (5–) 10–15 mm, tips thick and rigid;</span> <span class="statement" id="st-d0_s11" data-properties="anther coloration;anther coloration;anther coloration">anthers usually pink to purple;</span> <span class="statement" id="st-d0_s12" data-properties="nectar chamber some measurement">nectar chamber 5–11 mm.</span> <span class="statement" id="st-d0_s13" data-properties="fruit coloration;fruit coloration;fruit coloration;fruit coloration;fruit atypical some measurement;fruit some measurement"><b>Fruits </b>green to yellow-green or pink (rarely red), (15–) 20–35 mm, pulp white.</span> <span class="statement" id="st-d0_s14" data-properties="pulp coloration;2n chromosome count">2n = 22.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>unbranched or 1–12(–76)-branched, forming large mounds of branches to 300. <b>Stems</b> usually erect or nearly so, cylindric (spheric), (2–)5–70 × (3–)5–13 cm; ribs 5–8 or 8–12, crests slightly undulate (localized populations contain plants with strongly interrupted ribs); areoles 10–40 mm apart. <b>Spines</b> (0–)3–11 per areole, straight to curved or contorted, appressed (radial spines) or spreading to projecting outward (some radials and central spines when present), white to yellow, gray, or black; radial spines (0–)1–10 per areole, (0–)15–90 mm; central spines 0–1(–4) per areole, angular, (0–)50–120 mm. <b>Flowers</b> (4–)5–10 × 3–7 cm; flower tube 20–35 mm; flower tube hairs 1–2 mm; inner tepals bright orange-red to dark red, proximally paler (bases sometimes yellow or white), (18–)25–40 × (5–)10–15 mm, tips thick and rigid; anthers usually pink to purple; nectar chamber 5–11 mm. <b>Fruits</b> green to yellow-green or pink (rarely red), (15–)20–35 mm, pulp white. <b>2n</b> = 22.</span><!--
  
 
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|phenology=Flowering Apr–Jun; fruiting 2-2 1/2 months after flowering.
 
|habitat=Upper edge of Mojave Desert to coniferous forests, igneous and calcareous rock outcrops, cliffs, and sandy hills
 
|habitat=Upper edge of Mojave Desert to coniferous forests, igneous and calcareous rock outcrops, cliffs, and sandy hills
 
|elevation=1000-2800 m
 
|elevation=1000-2800 m
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|basionyms=
 
|basionyms=
 
|family=Cactaceae
 
|family=Cactaceae
 +
|phenology=Flowering Apr–Jun; fruiting 2-2 1/2 months after flowering.
 
|habitat=Upper edge of Mojave Desert to coniferous forests, igneous and calcareous rock outcrops, cliffs, and sandy hills
 
|habitat=Upper edge of Mojave Desert to coniferous forests, igneous and calcareous rock outcrops, cliffs, and sandy hills
 
|elevation=1000-2800 m
 
|elevation=1000-2800 m
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|publication year=1848
 
|publication year=1848
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_321.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V4/V4_321.xml
 
|subfamily=Cactaceae subfam. Cactoideae
 
|subfamily=Cactaceae subfam. Cactoideae
 
|genus=Echinocereus
 
|genus=Echinocereus
 
|species=Echinocereus triglochidiatus
 
|species=Echinocereus triglochidiatus
|2n chromosome count=22
 
|anther coloration=usually pink;purple
 
|areole arrangement=apart
 
|areole some measurement=10mm;40mm
 
|branch count=0;300
 
|central spine arrangement or shape=angular
 
|central spine atypical count=1;4
 
|central spine atypical some measurement=0mm;50mm
 
|central spine count=0;1
 
|central spine some measurement=50mm;120mm
 
|crest shape=undulate
 
|flower atypical length=4cm;5cm
 
|flower length=5cm;10cm
 
|flower tube hair some measurement=1mm;2mm
 
|flower tube some measurement=20mm;35mm
 
|flower width=3cm;7cm
 
|fruit atypical some measurement=15mm;20mm
 
|fruit coloration=green;yellow-green or pink
 
|fruit some measurement=20mm;35mm
 
|inner tepal atypical length=18mm;25mm
 
|inner tepal atypical width=5mm;10mm
 
|inner tepal coloration=paler;bright-orange-red;dark red
 
|inner tepal length=25mm;40mm
 
|inner tepal width=10mm;15mm
 
|mound size=large
 
|nectar chamber some measurement=5mm;11mm
 
|plant architecture=1-12(-76)-branched;unbranched
 
|pulp coloration=white
 
|rib count=8;12
 
|spine arrangement=radial
 
|spine arrangement or shape=contorted
 
|spine atypical count=0;1
 
|spine atypical some measurement=0mm;15mm
 
|spine coloration=white;yellow gray or black
 
|spine count=1;10
 
|spine course=straight;curved
 
|spine orientation=spreading;projecting
 
|spine some measurement=15mm;90mm
 
|stem atypical length=2cm;5cm
 
|stem atypical width=3cm;5cm
 
|stem length=5cm;70cm
 
|stem orientation=nearly;erect
 
|stem shape=cylindric
 
|stem width=5cm;13cm
 
|tip texture=rigid
 
|tip width=thick
 
 
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Revision as of 15:05, 27 July 2019

Plants unbranched or 1–12(–76)-branched, forming large mounds of branches to 300. Stems usually erect or nearly so, cylindric (spheric), (2–)5–70 × (3–)5–13 cm; ribs 5–8 or 8–12, crests slightly undulate (localized populations contain plants with strongly interrupted ribs); areoles 10–40 mm apart. Spines (0–)3–11 per areole, straight to curved or contorted, appressed (radial spines) or spreading to projecting outward (some radials and central spines when present), white to yellow, gray, or black; radial spines (0–)1–10 per areole, (0–)15–90 mm; central spines 0–1(–4) per areole, angular, (0–)50–120 mm. Flowers (4–)5–10 × 3–7 cm; flower tube 20–35 mm; flower tube hairs 1–2 mm; inner tepals bright orange-red to dark red, proximally paler (bases sometimes yellow or white), (18–)25–40 × (5–)10–15 mm, tips thick and rigid; anthers usually pink to purple; nectar chamber 5–11 mm. Fruits green to yellow-green or pink (rarely red), (15–)20–35 mm, pulp white. 2n = 22.


Phenology: Flowering Apr–Jun; fruiting 2-2 1/2 months after flowering.
Habitat: Upper edge of Mojave Desert to coniferous forests, igneous and calcareous rock outcrops, cliffs, and sandy hills
Elevation: 1000-2800 m

Distribution

V4 321-distribution-map.gif

Ariz., Calif., Colo., Nev., N.Mex., Utah.

Discussion

Echinocereus triglochidiatus is the earliest name for a large group of diploid and polyploid taxa treated as conspecific by L. D. Benson (1969, 1982). The tetraploids are now recognized separately as E. coccineus, including E. polyacanthus Engelmann of Mexico. The diploids in the flora area are here divided into two allopatric species: E. triglochidiatus and E. arizonicus.

Plants in the western portion of the range of Echinocereus triglochidiatus have been called E. triglochidiatus var. mojavensis (Engelmann & J. M. Bigelow) L. D. Benson. That taxon includes curly-spined plants (mainly in California) and straight-spined plants (including most populations in Arizona, Utah, and western Colorado). The latter were mapped by L. D. Benson (1969, 1982) as part of his concept of E. triglochidiatus var. melanacanthus. Plants with the fewest and largest spines, called E. triglochidiatus var. triglochidiatus, occupy the eastern portion of the species’ distribution. The largest spines, whether central or radial, of var. triglochidiatus are sharply angular in cross section and 1–2 mm thick.

Less distinctive plants have been called Echinocereus triglochidiatus var. gonacanthus, a name carelessly applied to miscellaneous plants throughout the range of var. triglochidiatus. A well-known population at White Sands, New Mexico, inhabits saline flats adjacent to pure gypsum dunes. The unusually large, southernmost plants at White Sands, New Mexico, shrink to the same size as northern plants when grown together in a common garden (D. Weniger 1970). The epithet inermis has been applied at various taxonomic ranks to individual plants with spines absent or nearly so in the eastern portion of var. mojavensis (in and around southeastern Utah).

A geographically distant tetraploid, Echinocereus coccineus var. paucispinus, superficially resembles some eastern E. triglochidiatus but may be distinguished by its relatively terete and more consistently straight spines. Small plants of var. triglochidiatus without reproductive structures might be confused with E. fendleri; records of E. triglochidatus from near the Mexican border probably are misidentifications of E. fendleri in vegetative condition.

Selected References

None.

Lower Taxa

None.
Allan D. Zimmerman +  and Bruce D. Parfitt +
Engelmann in F. A. Wislizenus +
Claret-cup cactus +  and kingcup cactus +
Ariz. +, Calif. +, Colo. +, Nev. +, N.Mex. +  and Utah. +
1000-2800 m +
Upper edge of Mojave Desert to coniferous forests, igneous and calcareous rock outcrops, cliffs, and sandy hills +
Flowering Apr–Jun +  and fruiting 2-2 1/2 months after flowering. +
in F. A. Wislizenus, Mem. Tour N. Mexico, +
Echinocereus triglochidiatus var. gonacanthus +, Echinocereus triglochidiatus var. inermis +  and Echinocereus triglochidiatus var. mojavensis +
Echinocereus triglochidiatus +
Echinocereus +
species +