Difference between revisions of "Gymnocarpium jessoense subsp. parvulum"

Sarvela

Ann. Bot. Fenn. 15: 103. 1978.

Common names: Nahanni oak fern gymnocarpe du japon sous-espèce fréle
Illustrated
Synonyms: Gymnocarpium continentale
Treatment appears in FNA Volume 2.
FNA>Volume Importer
 
(Added Nunavut to distribution; treatment was published before Nunavut split from N.W.T.)
 
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|common_names=Nahanni oak fern;gymnocarpe du japon sous-espèce fréle
 
|common_names=Nahanni oak fern;gymnocarpe du japon sous-espèce fréle
 +
|special_status={{Treatment/ID/Special_status
 +
|code=F
 +
|label=Illustrated
 +
}}
 
|basionyms=
 
|basionyms=
 
|synonyms={{Treatment/ID/Synonym
 
|synonyms={{Treatment/ID/Synonym
 
|name=Gymnocarpium continentale
 
|name=Gymnocarpium continentale
|authority=unknown
+
|authority=
 +
|rank=species
 
}}
 
}}
 
|hierarchy=Dryopteridaceae;Gymnocarpium;Gymnocarpium jessoense;Gymnocarpium jessoense subsp. parvulum
 
|hierarchy=Dryopteridaceae;Gymnocarpium;Gymnocarpium jessoense;Gymnocarpium jessoense subsp. parvulum
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}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="stem diameter"><b>Stems </b>0.5–1.5 mm diam.;</span> <span class="statement" id="st-d0_s1" data-properties="scale some measurement">scales 1–4 mm.</span> <span class="statement" id="st-d0_s2" data-properties="leaf reproduction;leaf some measurement"><b>Fertile </b>leaves usually 8–39 cm.</span> <span class="statement" id="st-d0_s3" data-properties="petiole some measurement;glandular-hair count"><b>Petiole </b>5–25 cm, with moderately abundant glandular-hairs distally;</span> <span class="statement" id="st-d0_s4" data-properties="scale some measurement">scales 2–6 mm.</span> <span class="statement" id="st-d0_s5" data-properties="blade shape;blade shape;blade shape;blade shape;blade some measurement;blade texture;blade fragility;blade architecture or arrangement;blade fragility;abaxial surface architecture or function or pubescence;rachis architecture or function or pubescence;adaxial surface pubescence"><b>Blade </b>narrowly deltate to narrowly ovate, 2-pinnate-pinnatifid, 3–14 cm, firm and robust or lax and delicate, abaxial surface moderately glandular, rachis moderately to densely glandular, adaxial surface glabrous.</span> <span class="statement" id="st-d0_s6" data-properties="pinna apex shape"><b>Pinna </b>apex acute.</span> <span class="statement" id="st-d0_s7" data-properties="proximal pinna some measurement;proximal pinna course;pinnule orientation;pinnule course"><b>Proximal </b>pinnae 2–9 cm, strongly curved toward apex of leaf, basiscopic pinnules strongly curved toward apex of pinna;</span> <span class="statement" id="st-d0_s8" data-properties="pinnule orientation;pinnule architecture;pinnule shape;pinnule shape;pinnule architecture;pinnulet orientation;pinnulet variability;pinnulet arrangement">basal basiscopic pinnule usually sessile, pinnatifid or rarely pinnate-pinnatifid, if sessile then with basal basiscopic pinnulet often equaling adjacent pinnulet;</span> <span class="statement" id="st-d0_s9" data-properties="pinnule orientation;pinnule architecture;pinnulet orientation;pinnulet variability;pinnulet arrangement">2d basal basiscopic pinnule sessile, with basal basiscopic pinnulet equaling adjacent pinnulet;</span> <span class="statement" id="st-d0_s10" data-properties="pinnule orientation;pinnule architecture;pinnulet orientation;pinnulet length or size;pinnulet variability;pinnulet arrangement">basal acroscopic pinnule sessile with basal basiscopic pinnulet longer than or equaling adjacent pinnulet.</span> <span class="statement" id="st-d0_s11" data-properties="pinna architecture;pinnule orientation;pinnule variability;pinnule height or length or size;pinnule arrangement"><b>Pinnae </b>of 2d pair almost always sessile with basal basiscopic pinnule usually equaling or slightly shorter than adjacent pinnule and equaling basal acroscopic pinnule;</span> <span class="statement" id="st-d0_s12" data-properties="pinnule orientation;pinnule variability;pinnule height or length or size;pinnule arrangement;apex architecture or shape;apex shape">basal acroscopic pinnule equaling or slightly shorter than adjacent pinnule, apex often entire, rounded.</span> <span class="statement" id="st-d0_s13" data-properties="pinna architecture;pinnule orientation;basal pinnule variability;basal pinnule arrangement;basal pinnule variability;basal pinnule orientation"><b>Pinnae </b>of 3d pair sessile with basal basiscopic pinnule equaling adjacent pinnule and equaling basal acroscopic pinnule;</span> <span class="statement" id="st-d0_s14" data-properties="pinnule orientation;pinnule variability;pinnule height or length or size;pinnule arrangement">basal acroscopic pinnule equaling or slightly shorter than adjacent pinnule.</span> <span class="statement" id="st-d0_s15" data-properties="ultimate segment shape;ultimate segment shape;ultimate segment shape;ultimate segment shape;apex architecture or shape;apex shape"><b>Ultimate </b>segments of proximal pinnae oblong, entire to slightly crenate, apex entire, rounded.</span> <span class="statement" id="st-d0_s16" data-properties="spore some measurement;2n chromosome count"><b>Spores </b>32–37 µm. <b>2n</b> = 160.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>0.5–1.5 mm diam.; scales 1–4 mm. <b>Fertile</b> leaves usually 8–39 cm. <b>Petiole</b> 5–25 cm, with moderately abundant glandular hairs distally; scales 2–6 mm. <b>Blade</b> narrowly deltate to narrowly ovate, 2-pinnate-pinnatifid, 3–14 cm, firm and robust or lax and delicate, abaxial surface moderately glandular, rachis moderately to densely glandular, adaxial surface glabrous. <b>Pinna</b> apex acute. <b>Proximal</b> pinnae 2–9 cm, strongly curved toward apex of leaf, basiscopic pinnules strongly curved toward apex of pinna; basal basiscopic pinnule usually sessile, pinnatifid or rarely pinnate-pinnatifid, if sessile then with basal basiscopic pinnulet often equaling adjacent pinnulet; 2d basal basiscopic pinnule sessile, with basal basiscopic pinnulet equaling adjacent pinnulet; basal acroscopic pinnule sessile with basal basiscopic pinnulet longer than or equaling adjacent pinnulet. <b>Pinnae</b> of 2d pair almost always sessile with basal basiscopic pinnule usually equaling or slightly shorter than adjacent pinnule and equaling basal acroscopic pinnule; basal acroscopic pinnule equaling or slightly shorter than adjacent pinnule, apex often entire, rounded. <b>Pinnae</b> of 3d pair sessile with basal basiscopic pinnule equaling adjacent pinnule and equaling basal acroscopic pinnule; basal acroscopic pinnule equaling or slightly shorter than adjacent pinnule. <b>Ultimate</b> segments of proximal pinnae oblong, entire to slightly crenate, apex entire, rounded. <b>Spores</b> 32–37 µm. <b>2n</b> = 160.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
 
|habitat=Acid or neutral substrates at summit of cool, shale talus slopes, and on granitic cliffs and outcrops
 
|habitat=Acid or neutral substrates at summit of cool, shale talus slopes, and on granitic cliffs and outcrops
 
|elevation=0–2000 m
 
|elevation=0–2000 m
|distribution=Alta.;B.C.;Man.;N.B.;N.W.T.;Ont.;Que.;Sask.;Yukon;Alaska;Conn.;Iowa;Maine;Mich.;Minn.;Vt.;Wis.;Europe in Finland;Asia in Siberia;Kazakhstan.
+
|distribution=Alta.;B.C.;Man.;N.B.;N.W.T.;Nunavut;Ont.;Que.;Sask.;Yukon;Alaska;Conn.;Iowa;Maine;Mich.;Minn.;Vt.;Wis.;Europe in Finland;Asia in Siberia;Kazakhstan.
|discussion=<p>Hybrids between Gymnocarpium jessoense subsp. parvulum and G. dryopteris (G. × intermedium Sarvela) are usually found wherever these two taxa occur together (Finland; Manitoba, Northwest Territories, Ontario, Quebec, Saskatchewan, Yukon; Alaska, Michigan, Minnesota, Wisconsin), and they are particularly abundant in the Great Lakes region. These hybrids have sometimes been referred to as G. × heterosporum, a name that is, however, correctly restricted to unique hybrids between G. robertianum and G. appalachianum (see discussion under G. robertianum; K. M. Pryer 1992). Gymnocarpium × intermedium is intermediate between the two parental species in its leaf morphology and glandularity, and it can be readily distinguished by its small, blackish, malformed, abortive spores, as well as large, brown, round spores that may allow this taxon to reproduce apogamously. Of the Gymnocarpium sterile hybrids, G. × intermedium is the easiest to distinguish morphologically.</p>
+
|discussion=<p>Hybrids between <i>Gymnocarpium jessoense </i>subsp.<i> parvulum</i> and <i>G. dryopteris</i> (G. × intermedium Sarvela) are usually found wherever these two taxa occur together (Finland; Manitoba, Northwest Territories, Ontario, Quebec, Saskatchewan, Yukon; Alaska, Michigan, Minnesota, Wisconsin), and they are particularly abundant in the Great Lakes region. These hybrids have sometimes been referred to as G. × heterosporum, a name that is, however, correctly restricted to unique hybrids between <i>G. robertianum</i> and <i>G. appalachianum</i> (see discussion under <i>G. robertianum</i>; K. M. Pryer 1992). <i>Gymnocarpium</i> × intermedium is intermediate between the two parental species in its leaf morphology and glandularity, and it can be readily distinguished by its small, blackish, malformed, abortive spores, as well as large, brown, round spores that may allow this taxon to reproduce apogamously. Of the <i>Gymnocarpium</i> sterile hybrids, G. × intermedium is the easiest to distinguish morphologically.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Gymnocarpium jessoense subsp. parvulum
 
name=Gymnocarpium jessoense subsp. parvulum
|author=
 
 
|authority=Sarvela
 
|authority=Sarvela
 
|rank=subspecies
 
|rank=subspecies
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|habitat=Acid or neutral substrates at summit of cool, shale talus slopes, and on granitic cliffs and outcrops
 
|habitat=Acid or neutral substrates at summit of cool, shale talus slopes, and on granitic cliffs and outcrops
 
|elevation=0–2000 m
 
|elevation=0–2000 m
|distribution=Alta.;B.C.;Man.;N.B.;N.W.T.;Ont.;Que.;Sask.;Yukon;Alaska;Conn.;Iowa;Maine;Mich.;Minn.;Vt.;Wis.;Europe in Finland;Asia in Siberia;Kazakhstan.
+
|distribution=Alta.;B.C.;Man.;N.B.;N.W.T.;Nunavut;Ont.;Que.;Sask.;Yukon;Alaska;Conn.;Iowa;Maine;Mich.;Minn.;Vt.;Wis.;Europe in Finland;Asia in Siberia;Kazakhstan.
 
|reference=None
 
|reference=None
 
|publication title=Ann. Bot. Fenn.
 
|publication title=Ann. Bot. Fenn.
 
|publication year=1978
 
|publication year=1978
|special status=
+
|special status=Illustrated
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_28.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_28.xml
 
|genus=Gymnocarpium
 
|genus=Gymnocarpium
 
|species=Gymnocarpium jessoense
 
|species=Gymnocarpium jessoense
 
|subspecies=Gymnocarpium jessoense subsp. parvulum
 
|subspecies=Gymnocarpium jessoense subsp. parvulum
|2n chromosome count=160
 
|abaxial surface architecture or function or pubescence=glandular
 
|adaxial surface pubescence=glabrous
 
|apex architecture or shape=entire;entire
 
|apex shape=rounded;rounded
 
|basal pinnule arrangement=adjacent
 
|basal pinnule orientation=acroscopic
 
|basal pinnule variability=equaling;equaling
 
|blade architecture or arrangement=lax
 
|blade fragility=delicate;robust
 
|blade shape=2-pinnate-pinnatifid;narrowly deltate;narrowly ovate
 
|blade some measurement=3cm;14cm
 
|blade texture=firm
 
|glandular-hair count=abundant
 
|leaf reproduction=fertile
 
|leaf some measurement=8cm;39cm
 
|petiole some measurement=5cm;25cm
 
|pinna apex shape=acute
 
|pinna architecture=sessile;sessile
 
|pinnule architecture=sessile;sessile;sessile;sessile
 
|pinnule arrangement=adjacent;adjacent;adjacent
 
|pinnule course=curved
 
|pinnule height or length or size=slightly shorter;slightly shorter;slightly shorter
 
|pinnule orientation=acroscopic;basiscopic;acroscopic;basiscopic;acroscopic;basiscopic;basiscopic;basiscopic
 
|pinnule shape=pinnate-pinnatifid;pinnatifid
 
|pinnule variability=equaling;equaling;equaling
 
|pinnulet arrangement=adjacent;adjacent;adjacent
 
|pinnulet length or size=longer
 
|pinnulet orientation=basiscopic;basiscopic;basiscopic
 
|pinnulet variability=equaling;equaling;equaling
 
|proximal pinna course=curved
 
|proximal pinna some measurement=2cm;9cm
 
|rachis architecture or function or pubescence=glandular
 
|scale some measurement=2mm;6mm
 
|spore some measurement=32um;37um
 
|stem diameter=0.5mm;1.5mm
 
|ultimate segment shape=entire;slightly crenate
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Gymnocarpium jessoense]]
 
-->[[Category:Treatment]][[Category:Gymnocarpium jessoense]]

Latest revision as of 17:37, 24 September 2021

Stems 0.5–1.5 mm diam.; scales 1–4 mm. Fertile leaves usually 8–39 cm. Petiole 5–25 cm, with moderately abundant glandular hairs distally; scales 2–6 mm. Blade narrowly deltate to narrowly ovate, 2-pinnate-pinnatifid, 3–14 cm, firm and robust or lax and delicate, abaxial surface moderately glandular, rachis moderately to densely glandular, adaxial surface glabrous. Pinna apex acute. Proximal pinnae 2–9 cm, strongly curved toward apex of leaf, basiscopic pinnules strongly curved toward apex of pinna; basal basiscopic pinnule usually sessile, pinnatifid or rarely pinnate-pinnatifid, if sessile then with basal basiscopic pinnulet often equaling adjacent pinnulet; 2d basal basiscopic pinnule sessile, with basal basiscopic pinnulet equaling adjacent pinnulet; basal acroscopic pinnule sessile with basal basiscopic pinnulet longer than or equaling adjacent pinnulet. Pinnae of 2d pair almost always sessile with basal basiscopic pinnule usually equaling or slightly shorter than adjacent pinnule and equaling basal acroscopic pinnule; basal acroscopic pinnule equaling or slightly shorter than adjacent pinnule, apex often entire, rounded. Pinnae of 3d pair sessile with basal basiscopic pinnule equaling adjacent pinnule and equaling basal acroscopic pinnule; basal acroscopic pinnule equaling or slightly shorter than adjacent pinnule. Ultimate segments of proximal pinnae oblong, entire to slightly crenate, apex entire, rounded. Spores 32–37 µm. 2n = 160.


Habitat: Acid or neutral substrates at summit of cool, shale talus slopes, and on granitic cliffs and outcrops
Elevation: 0–2000 m

Distribution

V2 28-distribution-map.gif

Alta., B.C., Man., N.B., N.W.T., Nunavut, Ont., Que., Sask., Yukon, Alaska, Conn., Iowa, Maine, Mich., Minn., Vt., Wis., Europe in Finland, Asia in Siberia, Kazakhstan.

Discussion

Hybrids between Gymnocarpium jessoense subsp. parvulum and G. dryopteris (G. × intermedium Sarvela) are usually found wherever these two taxa occur together (Finland; Manitoba, Northwest Territories, Ontario, Quebec, Saskatchewan, Yukon; Alaska, Michigan, Minnesota, Wisconsin), and they are particularly abundant in the Great Lakes region. These hybrids have sometimes been referred to as G. × heterosporum, a name that is, however, correctly restricted to unique hybrids between G. robertianum and G. appalachianum (see discussion under G. robertianum; K. M. Pryer 1992). Gymnocarpium × intermedium is intermediate between the two parental species in its leaf morphology and glandularity, and it can be readily distinguished by its small, blackish, malformed, abortive spores, as well as large, brown, round spores that may allow this taxon to reproduce apogamously. Of the Gymnocarpium sterile hybrids, G. × intermedium is the easiest to distinguish morphologically.

Selected References

None.

Lower Taxa

None.
Kathleen M. Pryer +
Sarvela +
Dryopteris jessoensis +
Nahanni oak fern +  and gymnocarpe du japon sous-espèce fréle +
Alta. +, B.C. +, Man. +, N.B. +, N.W.T. +, Nunavut +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Conn. +, Iowa +, Maine +, Mich. +, Minn. +, Vt. +, Wis. +, Europe in Finland +, Asia in Siberia +  and Kazakhstan. +
0–2000 m +
Acid or neutral substrates at summit of cool, shale talus slopes, and on granitic cliffs and outcrops +
Ann. Bot. Fenn. +
Illustrated +
Gymnocarpium continentale +
Gymnocarpium jessoense subsp. parvulum +
Gymnocarpium jessoense +
subspecies +