Difference between revisions of "Pinus torreyana"

Parry ex Carrière

326. 1855.

Common names: Torrey pine
EndemicConservation concern
Treatment appears in FNA Volume 2.
FNA>Volume Importer
 
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|common_names=Torrey pine
 
|common_names=Torrey pine
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|special_status={{Treatment/ID/Special_status
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|code=E
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|label=Endemic
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}}{{Treatment/ID/Special_status
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|code=C
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|label=Conservation concern
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}}
 
|basionyms=
 
|basionyms=
 
|synonyms=
 
|synonyms=
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--><span class="statement" id="st-d0_s0" data-properties="tree atypical some measurement;tree some measurement"><b>Trees </b>to 15 (–23) m;</span> <span class="statement" id="st-d0_s1" data-properties="trunk diameter;trunk orientation;nature course or orientation">trunk to 1m diam., in nature mostly crooked and leaning;</span> <span class="statement" id="st-d0_s2" data-properties="crown shape;crown shape;crown shape;crown architecture or course">crown rounded to flattened or irregular.</span> <span class="statement" id="st-d0_s3" data-properties="bark coloration;bark coloration;bark coloration;bark architecture;ridge architecture or course;ridge shape;ridge prominence or shape;ridge architecture or pubescence"><b>Bark </b>redbrown to purple-red, deeply furrowed with irregular, elongate, flat, scaly ridges.</span> <span class="statement" id="st-d0_s4" data-properties="branch architecture or course;branch orientation;branch shape"><b>Branches </b>irregular, spreading-ascending, candelabralike;</span> <span class="statement" id="st-d0_s5" data-properties="twig fragility or size;twig coloration;twig life cycle;twig depth;twig coloration;twig pubescence or relief">twigs stout (1–2cm thick), greenish, aging deep gray-brown to near black, rough.</span> <span class="statement" id="st-d0_s6" data-properties="bud shape;bud coloration;bud some measurement"><b>Buds </b>conic-ovoid, pale-brown, to 2.5cm;</span> <span class="statement" id="st-d0_s7" data-properties="scale margin shape">scale margins white-fringed.</span> <span class="statement" id="st-d0_s8" data-properties="leaf count;leaf orientation;leaf orientation;leaf duration;leaf count;leaf length;leaf width;leaf course;leaf course;leaf architecture;leaf reflectance;leaf coloration;stomatal width;line width;margin architecture or shape;apex shape"><b>Leaves </b>mostly 5 per fascicle, ascending or spreading, persisting 3–4 years, 15–30cm × ca. 2mm, straight or curved, slightly twisted, dull gray-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly acute;</span> <span class="statement" id="st-d0_s9" data-properties="sheath some measurement;sheath duration">sheath to 2cm, shed early, base persistent.</span> <span class="statement" id="st-d0_s10" data-properties="pollen cone shape;pollen cone some measurement;pollen cone coloration"><b>Pollen </b>cones ovoid, 20–30mm, yellow.</span> <span class="statement" id="st-d0_s11" data-properties="seed-cone life cycle;seed-cone duration;seed-cone position;seed-cone size;seed-cone architecture or shape;seed-cone shape;seed-cone shape;seed-cone distance;seed-cone coloration;seed-cone coloration;seed-cone coloration;seed-cone reflectance;year count;year count;stalk some measurement"><b>Seed-</b>cones maturing in 3 years, shedding seeds soon thereafter, persisting to 5 years, lateral, massive, heavy, symmetric, ovoid before opening, broadly ovoid when open, 10–15cm, yellow to redbrown, lustrous, stalks to 4cm;</span> <span class="statement" id="st-d0_s12" data-properties="apophysis width;apophysis shape;keel count;keel position;keel arrangement">apophyses thick, angulately dome-shaped, with 5 low convergent keels;</span> <span class="statement" id="st-d0_s13" data-properties="umbo position;pyramid height or length or size;pyramid architecture">umbo central, forming short, curved-tipped pyramid.</span> <span class="statement" id="st-d0_s14" data-properties="seed shape"><b>Seeds </b>narrowly obovoid;</span> <span class="statement" id="st-d0_s15" data-properties="body some measurement;body coloration;body coloration">body 16–24mm, brown, apically dark-brown;</span> <span class="statement" id="st-d0_s16" data-properties="wing width;wing architecture;wing some measurement">wing broad, oblique-tipped, to 15mm.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Trees </b>to 15(–23)m; trunk to 1m diam., in nature mostly crooked and leaning; crown rounded to flattened or irregular. <b>Bark</b> red-brown to purple-red, deeply furrowed with irregular, elongate, flat, scaly ridges. <b>Branches</b> irregular, spreading-ascending, candelabralike; twigs stout (1–2cm thick), greenish, aging deep gray-brown to near black, rough. <b>Buds</b> conic-ovoid, pale brown, to 2.5cm; scale margins white-fringed. <b>Leaves</b> mostly 5 per fascicle, ascending or spreading, persisting 3–4 years, 15–30cm × ca. 2mm, straight or curved, slightly twisted, dull gray-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly acute; sheath to 2cm, shed early, base persistent. <b>Pollen</b> cones ovoid, 20–30mm, yellow. <b>Seed</b> cones maturing in 3 years, shedding seeds soon thereafter, persisting to 5 years, lateral, massive, heavy, symmetric, ovoid before opening, broadly ovoid when open, 10–15cm, yellow- to red-brown, lustrous, stalks to 4cm; apophyses thick, angulately dome-shaped, with 5 low convergent keels; umbo central, forming short, curved-tipped pyramid. <b>Seeds</b> narrowly obovoid; body 16–24mm, brown, apically dark brown; wing broad, oblique-tipped, to 15mm.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
|distribution=Only in the flora.
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|distribution=Calif.;only in the flora.
|discussion=<p>Pinus torreyana is a rare and local Tertiary relic species whose present range is reduced to two small areas of southern California: near Del Mar (San Diego County) and on the northeastern shore of Santa Rosa Island (Santa Barbara County). Its distribution in Oligocene and Miocene (or at least that of its near ancestor) extended north to Oregon. Its harsh natural habitat elicits an unusually contorted and often sparse form, quite unlike the cleaner and taller form the species takes in cultivation.</p><!--
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|discussion=<p><i>Pinus torreyana</i> is a rare and local Tertiary relic species whose present range is reduced to two small areas of southern California: near Del Mar (San Diego County) and on the northeastern shore of Santa <i>Rosa</i> Island (Santa Barbara County). Its distribution in Oligocene and Miocene (or at least that of its near ancestor) extended north to Oregon. Its harsh natural habitat elicits an unusually contorted and often sparse form, quite unlike the cleaner and taller form the species takes in cultivation.</p><!--
--><p>In terms of numbers of individuals in the wild, as well as the small area occupied by natural populations, Pinus torreyana is without a doubt the rarest North American pine. As such it is under protection. Artificial crosses between it and another, more widespread Tertiary relic, P. sabiniana, have been successful.</p><!--
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--><p>In terms of numbers of individuals in the wild, as well as the small area occupied by natural populations, <i>Pinus torreyana</i> is without a doubt the rarest North American pine. As such it is under protection. Artificial crosses between it and another, more widespread Tertiary relic, <i>P. sabiniana</i>, have been successful.</p><!--
 
--><p>Subspecies 2.</p>
 
--><p>Subspecies 2.</p>
 
|tables=
 
|tables=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Pinus torreyana
 
name=Pinus torreyana
|author=
 
 
|authority=Parry ex Carrière
 
|authority=Parry ex Carrière
 
|rank=species
 
|rank=species
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|basionyms=
 
|basionyms=
 
|family=Pinaceae
 
|family=Pinaceae
|distribution=Only in the flora.
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|distribution=Calif.;only in the flora.
 
|reference=haller1986a
 
|reference=haller1986a
 
|publication title=
 
|publication title=
 
|publication year=1855
 
|publication year=1855
|special status=
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|special status=Endemic;Conservation concern
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_140.xml
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|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_140.xml
 
|genus=Pinus
 
|genus=Pinus
 
|species=Pinus torreyana
 
|species=Pinus torreyana
|apex shape=acute
 
|apophysis shape=dome-shaped
 
|apophysis width=thick
 
|bark architecture=furrowed
 
|bark coloration=redbrown;purple-red
 
|body coloration=dark-brown;brown
 
|body some measurement=16mm;24mm
 
|branch architecture or course=irregular
 
|branch orientation=spreading-ascending
 
|branch shape=candelabralike
 
|bud coloration=pale-brown
 
|bud shape=conic-ovoid
 
|bud some measurement=0cm;2.5cm
 
|crown architecture or course=irregular
 
|crown shape=rounded;flattened
 
|keel arrangement=convergent
 
|keel count=5
 
|keel position=low
 
|leaf architecture=twisted
 
|leaf coloration=gray-green
 
|leaf count=3;4
 
|leaf course=curved;straight
 
|leaf duration=persisting
 
|leaf length=15cm;30cm
 
|leaf orientation=spreading;ascending
 
|leaf reflectance=dull
 
|leaf width=2
 
|line width=fine
 
|margin architecture or shape=serrulate
 
|nature course or orientation=crooked
 
|pollen cone coloration=yellow
 
|pollen cone shape=ovoid
 
|pollen cone some measurement=20mm;30mm
 
|pyramid architecture=curved-tipped
 
|pyramid height or length or size=short
 
|ridge architecture or course=irregular
 
|ridge architecture or pubescence=scaly
 
|ridge prominence or shape=flat
 
|ridge shape=elongate
 
|scale margin shape=white-fringed
 
|seed shape=obovoid
 
|seed-cone architecture or shape=symmetric
 
|seed-cone coloration=yellow;redbrown
 
|seed-cone distance=10cm;15cm
 
|seed-cone duration=persisting
 
|seed-cone life cycle=maturing
 
|seed-cone position=lateral
 
|seed-cone reflectance=lustrous
 
|seed-cone shape=ovoid;ovoid
 
|seed-cone size=massive
 
|sheath duration=persistent
 
|sheath some measurement=0cm;2cm
 
|stalk some measurement=4
 
|stomatal width=fine
 
|tree atypical some measurement=15m;23m
 
|tree some measurement=0m;15m
 
|trunk diameter=0m;1m
 
|trunk orientation=leaning
 
|twig coloration=gray-brown;greenish
 
|twig depth=deep
 
|twig fragility or size=stout
 
|twig life cycle=aging
 
|twig pubescence or relief=rough
 
|umbo position=central
 
|wing architecture=oblique-tipped
 
|wing some measurement=0mm;15mm
 
|wing width=broad
 
|year count=5;3
 
 
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[[Category:Treatment]]
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[[Category:Pinus]]
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[[Category:Revised Since Print]]

Latest revision as of 17:56, 6 November 2020

Trees to 15(–23)m; trunk to 1m diam., in nature mostly crooked and leaning; crown rounded to flattened or irregular. Bark red-brown to purple-red, deeply furrowed with irregular, elongate, flat, scaly ridges. Branches irregular, spreading-ascending, candelabralike; twigs stout (1–2cm thick), greenish, aging deep gray-brown to near black, rough. Buds conic-ovoid, pale brown, to 2.5cm; scale margins white-fringed. Leaves mostly 5 per fascicle, ascending or spreading, persisting 3–4 years, 15–30cm × ca. 2mm, straight or curved, slightly twisted, dull gray-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly acute; sheath to 2cm, shed early, base persistent. Pollen cones ovoid, 20–30mm, yellow. Seed cones maturing in 3 years, shedding seeds soon thereafter, persisting to 5 years, lateral, massive, heavy, symmetric, ovoid before opening, broadly ovoid when open, 10–15cm, yellow- to red-brown, lustrous, stalks to 4cm; apophyses thick, angulately dome-shaped, with 5 low convergent keels; umbo central, forming short, curved-tipped pyramid. Seeds narrowly obovoid; body 16–24mm, brown, apically dark brown; wing broad, oblique-tipped, to 15mm.

Discussion

Pinus torreyana is a rare and local Tertiary relic species whose present range is reduced to two small areas of southern California: near Del Mar (San Diego County) and on the northeastern shore of Santa Rosa Island (Santa Barbara County). Its distribution in Oligocene and Miocene (or at least that of its near ancestor) extended north to Oregon. Its harsh natural habitat elicits an unusually contorted and often sparse form, quite unlike the cleaner and taller form the species takes in cultivation.

In terms of numbers of individuals in the wild, as well as the small area occupied by natural populations, Pinus torreyana is without a doubt the rarest North American pine. As such it is under protection. Artificial crosses between it and another, more widespread Tertiary relic, P. sabiniana, have been successful.

Subspecies 2.

Key

1 Crown of sheltered tree narrower than height of mature tree, open with well-spaced branches in natural sites; seed cones less than 13.5 cm broad, as broad as long or longer; only tip of umbo curved outward; maximum seed width ca. 12 mm. Pinus torreyana subsp. torreyana
1 Crown of sheltered tree broader than height of mature tree, compact in natural sites; seed cones mostly more than 13.5 cm broad, broader than long; entire umbo curved outward; maximum seed width ca. 14 mm. Pinus torreyana subsp. insularis