Difference between revisions of "Sclerocactus sileri"

(L. D. Benson) K. D. Heil & J. M. Porter

Haseltonia 2: 39. 1994.

Common names: Siler fishhook cactus
Basionym: Sclerocactus pubispinus var. sileri L. D. Benson
Synonyms: Pediocactus pubispinus subsp. sileri (L. D. Benson) HaldaPediocactus whipplei subsp. busekii (Hochstätter) HaldaSclerocactus whipplei subsp. busekii Hochstätter
Treatment appears in FNA Volume 4. Treatment on page 205. Mentioned on page 198, 199.
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--><span class="statement" id="st-d0_s0" data-properties="stem architecture;stem coloration;stem shape;stem shape;stem shape;stem shape;stem length;stem width"><b>Stems </b>unbranched (2-branched), green, depressed-spheric to short cylindric, 4–8.5 × 3–5.5 cm;</span> <span class="statement" id="st-d0_s1" data-properties="rib atypical count;rib atypical count;rib count;rib development;tubercle prominence;rib position">ribs usually (12–) 13 (–14), not well developed, tubercles prominent on low ribs.</span> <span class="statement" id="st-d0_s2" data-properties=""><b>Spines </b>obscuring stems or nearly so;</span> <span class="statement" id="st-d0_s3" data-properties="spine arrangement;spine atypical count;spine count;spine shape;spine arrangement or shape;spine length;spine width">radial spines 6–8 (–12) per areole, acicular, elliptic in cross-section, 5–15 × 0.5–0.8 mm;</span> <span class="statement" id="st-d0_s4" data-properties="central spine count">central spines 4–5 per areole;</span> <span class="statement" id="st-d0_s5" data-properties="abaxial central spine count;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine orientation;abaxial central spine course;abaxial central spine orientation;abaxial central spine shape;abaxial central spine shape;abaxial central spine shape;abaxial central spine shape;abaxial central spine shape;abaxial central spine shape;abaxial central spine length;abaxial central spine width">abaxial central spines 1–2 per areole, white, gray, or purplish tinged, turned or curving somewhat downward, angled, strongly hooked, slightly contorted, terete to angled, 12–35 × 0.5–1 mm;</span> <span class="statement" id="st-d0_s6" data-properties="lateral central spine count;lateral central spine shape;abaxial spine height or length or size">lateral central spines 2 per areole, similar to abaxial but slightly shorter, usually not hooked;</span> <span class="statement" id="st-d0_s7" data-properties="adaxial central spine count;adaxial central spine orientation;adaxial central spine orientation;adaxial central spine coloration;adaxial central spine coloration;adaxial central spine coloration;adaxial central spine shape;adaxial central spine prominence;adaxial central spine length;adaxial central spine width">adaxial central spines 1–2 per areole, erect, recurved, white to tan, strongly flattened, conspicuous, 13–40 × 0.8–2 mm.</span> <span class="statement" id="st-d0_s8" data-properties="flower shape;flower shape;flower shape;flower length;flower width"><b>Flowers </b>funnelform to campanulate, 2.5–3 × 2–3 cm;</span> <span class="statement" id="st-d0_s9" data-properties="outer tepal size;outer tepal shape;outer tepal length;outer tepal width;margin coloration;margin coloration;apex shape;margin texture;margin shape;margin shape">outer tepals with brownish and yellowish margins, larger oblanceolate, 10–17 × 5–8 mm, apex mucronate, margins membranous, crisped or minutely toothed;</span> <span class="statement" id="st-d0_s10" data-properties="inner tepal coloration;inner tepal coloration;largest tepal shape;largest tepal length;largest tepal width;apex shape">inner tepals yellow, sometimes suffused with brown, largest tepals oblanceolate, 15–25 × 5–7 mm, apex mucronate;</span> <span class="statement" id="st-d0_s11" data-properties="filament coloration;filament coloration;filament coloration">filaments white to greenish white;</span> <span class="statement" id="st-d0_s12" data-properties="anther coloration">anthers yellow;</span> <span class="statement" id="st-d0_s13" data-properties="fruit relief;fruit dehiscence;fruit coloration;fruit coloration;fruit coloration;fruit coloration;fruit shape;fruit length;fruit atypical width;fruit width;fruit condition or texture;slit count;slit orientation">styles minutely but densely papillate Fruits dehiscent along 2–4 vertical slits, green, turning red to tan, ovoid, 8–25 × 8–14 (–22) mm, dry;</span> <span class="statement" id="st-d0_s14" data-properties="scale count;scale texture;scale architecture;scale shape;scale shape">scales few, membranous, scarious-margined, minutely toothed or fringed.</span> <span class="statement" id="st-d0_s15" data-properties="seed coloration;seed coloration;seed coloration;seed length;seed width"><b>Seeds </b>black to light-brown, 2.7–3 × 2.2–2.5 mm;</span> <span class="statement" id="st-d0_s16" data-properties="papilla shape">testa with rounded papillae.</span><!--
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--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>unbranched (2-branched), green, depressed-spheric to short cylindric, 4–8.5 × 3–5.5 cm; ribs usually (12–)13(–14), not well developed, tubercles prominent on low ribs. <b>Spines</b> obscuring stems or nearly so; radial spines 6–8(–12) per areole, acicular, elliptic in cross section, 5–15 × 0.5–0.8 mm; central spines 4–5 per areole; abaxial central spines 1–2 per areole, white, gray, or purplish tinged, turned or curving somewhat downward, angled, strongly hooked, slightly contorted, terete to angled, 12–35 × 0.5–1 mm; lateral central spines 2 per areole, similar to abaxial but slightly shorter, usually not hooked; adaxial central spines 1–2 per areole, erect, recurved, white to tan, strongly flattened, conspicuous, 13–40 × 0.8–2 mm. <b>Flowers</b> funnelform to campanulate, 2.5–3 × 2–3 cm; outer tepals with brownish and yellowish margins, larger oblanceolate, 10–17 × 5–8 mm, apex mucronate, margins membranous, crisped or minutely toothed; inner tepals yellow, sometimes suffused with brown, largest tepals oblanceolate, 15–25 × 5–7 mm, apex mucronate; filaments white to greenish white; anthers yellow; styles minutely but densely papillate Fruits dehiscent along 2–4 vertical slits, green, turning red to tan, ovoid, 8–25 × 8–14(–22) mm, dry; scales few, membranous, scarious-margined, minutely toothed or fringed. <b>Seeds</b> black to light brown, 2.7–3 × 2.2–2.5 mm; testa with rounded papillae.</span><!--
  
 
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|phenology=Flowering late Apr–May.
 
|habitat=Silty sand or clay soils, often with gravel, grama grasslands with dropseed, yucca, hedgehog cactus, sagebrush, and snakeweed, pinyon-juniper woodlands
 
|habitat=Silty sand or clay soils, often with gravel, grama grasslands with dropseed, yucca, hedgehog cactus, sagebrush, and snakeweed, pinyon-juniper woodlands
 
|elevation=1600-1700 m
 
|elevation=1600-1700 m
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|basionyms=Sclerocactus pubispinus var. sileri
 
|basionyms=Sclerocactus pubispinus var. sileri
 
|family=Cactaceae
 
|family=Cactaceae
 +
|phenology=Flowering late Apr–May.
 
|habitat=Silty sand or clay soils, often with gravel, grama grasslands with dropseed, yucca, hedgehog cactus, sagebrush, and snakeweed, pinyon-juniper woodlands
 
|habitat=Silty sand or clay soils, often with gravel, grama grasslands with dropseed, yucca, hedgehog cactus, sagebrush, and snakeweed, pinyon-juniper woodlands
 
|elevation=1600-1700 m
 
|elevation=1600-1700 m
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|publication year=1994
 
|publication year=1994
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_383.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V4/V4_383.xml
 
|subfamily=Cactaceae subfam. Cactoideae
 
|subfamily=Cactaceae subfam. Cactoideae
 
|genus=Sclerocactus
 
|genus=Sclerocactus
 
|species=Sclerocactus sileri
 
|species=Sclerocactus sileri
|abaxial central spine coloration=purplish tinged;gray;purplish tinged;gray;white
 
|abaxial central spine count=1;2
 
|abaxial central spine course=curving
 
|abaxial central spine length=12mm;35mm
 
|abaxial central spine orientation=downward;turned
 
|abaxial central spine shape=terete;angled
 
|abaxial central spine width=0.5mm;1mm
 
|abaxial spine height or length or size=shorter
 
|adaxial central spine coloration=white;tan
 
|adaxial central spine count=1;2
 
|adaxial central spine length=13mm;40mm
 
|adaxial central spine orientation=recurved;erect
 
|adaxial central spine prominence=conspicuous
 
|adaxial central spine shape=flattened
 
|adaxial central spine width=0.8mm;2mm
 
|anther coloration=yellow
 
|apex shape=mucronate;mucronate
 
|central spine count=4;5
 
|filament coloration=white;greenish white
 
|flower length=2.5cm;3cm
 
|flower shape=funnelform;campanulate
 
|flower width=2cm;3cm
 
|fruit atypical width=14mm;22mm
 
|fruit coloration=red;tan
 
|fruit condition or texture=dry
 
|fruit dehiscence=dehiscent
 
|fruit length=8mm;25mm
 
|fruit relief=papillate
 
|fruit shape=ovoid
 
|fruit width=8mm;14mm
 
|inner tepal coloration=suffused with brown;yellow
 
|largest tepal length=15mm;25mm
 
|largest tepal shape=oblanceolate
 
|largest tepal width=5mm;7mm
 
|lateral central spine count=2
 
|lateral central spine shape=hooked
 
|margin coloration=yellowish;brownish
 
|margin shape=toothed;crisped
 
|margin texture=membranous
 
|outer tepal length=10mm;17mm
 
|outer tepal shape=oblanceolate
 
|outer tepal size=larger
 
|outer tepal width=5mm;8mm
 
|papilla shape=rounded
 
|rib atypical count=13;14
 
|rib count=13
 
|rib development=developed
 
|rib position=low
 
|scale architecture=scarious-margined
 
|scale count=few
 
|scale shape=fringed;toothed
 
|scale texture=membranous
 
|seed coloration=black;light-brown
 
|seed length=2.7mm;3mm
 
|seed width=2.2mm;2.5mm
 
|slit count=2;4
 
|slit orientation=vertical
 
|spine arrangement=radial
 
|spine arrangement or shape=elliptic
 
|spine atypical count=8;12
 
|spine count=6;8
 
|spine length=5mm;15mm
 
|spine shape=acicular
 
|spine width=0.5mm;0.8mm
 
|stem architecture=unbranched
 
|stem coloration=green
 
|stem length=4cm;8.5cm
 
|stem shape=depressed-spheric;short cylindric
 
|stem width=3cm;5.5cm
 
|tubercle prominence=prominent
 
 
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Revision as of 15:08, 27 July 2019

Stems unbranched (2-branched), green, depressed-spheric to short cylindric, 4–8.5 × 3–5.5 cm; ribs usually (12–)13(–14), not well developed, tubercles prominent on low ribs. Spines obscuring stems or nearly so; radial spines 6–8(–12) per areole, acicular, elliptic in cross section, 5–15 × 0.5–0.8 mm; central spines 4–5 per areole; abaxial central spines 1–2 per areole, white, gray, or purplish tinged, turned or curving somewhat downward, angled, strongly hooked, slightly contorted, terete to angled, 12–35 × 0.5–1 mm; lateral central spines 2 per areole, similar to abaxial but slightly shorter, usually not hooked; adaxial central spines 1–2 per areole, erect, recurved, white to tan, strongly flattened, conspicuous, 13–40 × 0.8–2 mm. Flowers funnelform to campanulate, 2.5–3 × 2–3 cm; outer tepals with brownish and yellowish margins, larger oblanceolate, 10–17 × 5–8 mm, apex mucronate, margins membranous, crisped or minutely toothed; inner tepals yellow, sometimes suffused with brown, largest tepals oblanceolate, 15–25 × 5–7 mm, apex mucronate; filaments white to greenish white; anthers yellow; styles minutely but densely papillate Fruits dehiscent along 2–4 vertical slits, green, turning red to tan, ovoid, 8–25 × 8–14(–22) mm, dry; scales few, membranous, scarious-margined, minutely toothed or fringed. Seeds black to light brown, 2.7–3 × 2.2–2.5 mm; testa with rounded papillae.


Phenology: Flowering late Apr–May.
Habitat: Silty sand or clay soils, often with gravel, grama grasslands with dropseed, yucca, hedgehog cactus, sagebrush, and snakeweed, pinyon-juniper woodlands
Elevation: 1600-1700 m

Discussion

Of conservation concern.

As far as is known, based on field investigations and herbarium collections, Sclerocactus sileri is restricted to northeast Coconino County, Arizona. This enigmatic cactus shares morphologic characteristics with both Sclerocactus whipplei (F. Hochstätter 1996) and S. pubispinus. The floral morphology and color is reminiscent of S. whipplei; however, the adaxial central spine morphology and fruit dehiscence is much like that found in S. pubispinus and S. spinosior. Phylogenetic analyses of chloroplast DNA sequences (J. M. Porter et al. 2000) provided evidence that S. sileri is the sister taxon to a clade including S. brevispinus, S. whipplei, S. cloverae, S. glaucus, S. parviflorus, S. wetlandicus, and S. wrightiae. Therefore, there is no support for immediate links between S. sileri and either S. whipplei or S. pubispinus.

Selected References

None.

Lower Taxa

None.
... more about "Sclerocactus sileri"
Kenneth D. Heil +  and J. Mark Porter +
(L. D. Benson) K. D. Heil & J. M. Porter +
Sclerocactus pubispinus var. sileri +
Siler fishhook cactus +
1600-1700 m +
Silty sand or clay soils, often with gravel, grama grasslands with dropseed, yucca, hedgehog cactus, sagebrush, and snakeweed, pinyon-juniper woodlands +
Flowering late Apr–May. +
Pediocactus pubispinus subsp. sileri +, Pediocactus whipplei subsp. busekii +  and Sclerocactus whipplei subsp. busekii +
Sclerocactus sileri +
Sclerocactus +
species +