Difference between revisions of "Selaginella mutica"

D. C. Eaton ex L. Underwood

Bull. Torrey Bot. Club 25: 128. 1898.

IllustratedEndemic
Treatment appears in FNA Volume 2.
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|place=25: 128. 1898
 
|place=25: 128. 1898
 
|year=1898
 
|year=1898
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|special_status={{Treatment/ID/Special_status
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|code=F
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|label=Illustrated
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|code=E
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|label=Endemic
 
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|basionyms=
 
|basionyms=
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--><span class="statement" id="st-d0_s0" data-properties="mat architecture or fragility"><b>Plants </b>on rock or terrestrial, forming loose mats.</span> <span class="statement" id="st-d0_s1" data-properties="stem architecture or shape;stem length or size;stem growth form or orientation;stem shape;branch architecture or shape;tip course"><b>Stems </b>radially symmetric, long to short-creeping, not readily fragmenting, ± regularly forked, without budlike arrested branches, tips straight;</span> <span class="statement" id="st-d0_s2" data-properties="main-stem development;lateral branch development;lateral branch shape">main-stem indeterminate, lateral branches determinate, 1–2-forked.</span> <span class="statement" id="st-d0_s3" data-properties="rhizophore diameter"><b>Rhizophores </b>borne on upperside of stems, throughout stem length, 0.13–0.23 mm diam.</span> <span class="statement" id="st-d0_s4" data-properties="leaf architecture;leaf fixation or orientation;leaf orientation;leaf coloration;leaf shape;leaf shape;leaf shape;leaf shape;leaf length;leaf width;pseudowhorl arrangement;pseudowhorl count"><b>Leaves </b>monomorphic, in ± alternate pseudowhorls of 3, tightly appressed, ascending, green, lanceolate to linear-lanceolate or lanceolate-elliptic, 1–2 X 0.45–0.6 mm;</span> <span class="statement" id="st-d0_s5" data-properties="abaxial ridge prominence">abaxial ridges well defined;</span> <span class="statement" id="st-d0_s6" data-properties="base shape;base fusion;base shape;base pubescence;base pubescence">base rounded and adnate, sometimes slightly decurrent, pubescent or glabrous;</span> <span class="statement" id="st-d0_s7" data-properties="margin shape;margin shape;margin shape;cilium coloration;cilium orientation;cilium orientation;cilium some measurement">margins ciliate to denticulate, cilia transparent, spreading or ascending, 0.03–0.17 mm;</span> <span class="statement" id="st-d0_s8" data-properties="apex shape;apex shape;apex shape;apex architecture or shape;apex shape;apex shape;apex shape">apex keeled, obtuse or slightly attenuate, nearly truncate in profile, blunt to short-bristled;</span> <span class="statement" id="st-d0_s9" data-properties="bristle coloration;bristle coloration;bristle coloration;bristle coloration;bristle architecture or pubescence or relief;bristle some measurement">bristle transparent to greenish transparent or whitish, smooth, 0.06–0.45 mm.</span> <span class="statement" id="st-d0_s10" data-properties="strobilus architecture or arrangement or growth form;strobilus atypical some measurement;strobilus some measurement"><b>Strobili </b>solitary, (0.6–) 1–3 cm;</span> <span class="statement" id="st-d0_s11" data-properties="sporophyll shape;sporophyll shape;sporophyll shape;sporophyll shape;sporophyll shape;abaxial ridge prominence;base pubescence;margin shape;margin shape;margin shape">sporophylls ovatelanceolate, ovate-elliptic, or deltate-ovate, abaxial ridges well defined, base glabrous, margins ciliate to denticulate, apex strongly to slightly keeled, short-bristled to blunt.</span> <span class="statement" id="st-d0_s12" data-properties="apex shape;apex shape;apex shape;apex shape;2n chromosome count">2n = 18.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>on rock or terrestrial, forming loose mats. <b>Stems</b> radially symmetric, long- to short-creeping, not readily fragmenting, ± regularly forked, without budlike arrested branches, tips straight; main stem indeterminate, lateral branches determinate, 1–2-forked. <b>Rhizophores</b> borne on upperside of stems, throughout stem length, 0.13–0.23 mm diam. <b>Leaves</b> monomorphic, in ± alternate pseudowhorls of 3, tightly appressed, ascending, green, lanceolate to linear-lanceolate or lanceolate-elliptic, 1–2 × 0.45–0.6 mm; abaxial ridges well defined; base rounded and adnate, sometimes slightly decurrent, pubescent or glabrous; margins ciliate to denticulate, cilia transparent, spreading or ascending, 0.03–0.17 mm; apex keeled, obtuse or slightly attenuate, nearly truncate in profile, blunt to short-bristled; bristle transparent to greenish transparent or whitish, smooth, 0.06–0.45 mm. <b>Strobili</b> solitary, (0.6–)1–3 cm; sporophylls ovate-lanceolate, ovate-elliptic, or deltate-ovate, abaxial ridges well defined, base glabrous, margins ciliate to denticulate, apex strongly to slightly keeled, short-bristled to blunt. <b>2n</b> = 18.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
|distribution=Only in the flora.
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|distribution=Ariz.;Colo.;N.Mex.;Tex.;Utah;Wyo.;only in the flora.
|discussion=<p>Selaginella mutica, S. underwoodii (R. M. Tryon 1955; C. A. Weatherby 1943), and S. wallacei all have similar patterns of variation. Study is needed to assess to what degree such variability is caused by environmental or genetic factors. Within S. mutica, two rather distinct, morphologic extremes are recognized here as varieties. Many specimens having leaves with spreading, long, marginal cilia and a short, broken, apical bristle have been considered intermediate between the two varieties, but they belong in S. mutica var. mutica.</p><!--
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|discussion=<p><i>Selaginella mutica</i>, <i>S. underwoodii</i> (R. M. Tryon 1955; C. A. Weatherby 1943), and <i>S. wallacei</i> all have similar patterns of variation. Study is needed to assess to what degree such variability is caused by environmental or genetic factors. Within <i>S. mutica</i>, two rather distinct, morphologic extremes are recognized here as varieties. Many specimens having leaves with spreading, long, marginal cilia and a short, broken, apical bristle have been considered intermediate between the two varieties, but they belong in <i>S. mutica </i>var.<i> mutica</i>.</p><!--
--><p>Selaginella mutica may be one of the parent species of the putative hybrid species S. × neomexicana (see discussion). Selaginella mutica is often found growing in the same habitat with S. underwoodii, S. × neomexicana, and S. weatherbiana. According to R. M. Tryon (1955), where the two grow together, S. mutica mats gradually entirely replace mats of S. underwoodii over time. Selaginella mutica is sometimes confused with S. viridissima.</p><!--
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--><p><i>Selaginella mutica</i> may be one of the parent species of the putative hybrid species <i>S.</i> × <i>neomexicana</i> (see discussion). <i>Selaginella mutica</i> is often found growing in the same habitat with <i>S. underwoodii</i>, <i>S.</i> × <i>neomexicana</i>, and <i>S. weatherbiana</i>. According to R. M. Tryon (1955), where the two grow together, <i>S. mutica</i> mats gradually entirely replace mats of <i>S. underwoodii</i> over time. <i>Selaginella mutica</i> is sometimes confused with <i>S. viridissima</i>.</p><!--
 
--><p>Varieties 2.</p>
 
--><p>Varieties 2.</p>
 
|tables=
 
|tables=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Selaginella mutica
 
name=Selaginella mutica
|author=
 
 
|authority=D. C. Eaton ex L. Underwood
 
|authority=D. C. Eaton ex L. Underwood
 
|rank=species
 
|rank=species
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|basionyms=
 
|basionyms=
 
|family=Selaginellaceae
 
|family=Selaginellaceae
|distribution=Only in the flora.
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|distribution=Ariz.;Colo.;N.Mex.;Tex.;Utah;Wyo.;only in the flora.
 
|reference=None
 
|reference=None
 
|publication title=Bull. Torrey Bot. Club
 
|publication title=Bull. Torrey Bot. Club
 
|publication year=1898
 
|publication year=1898
|special status=
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|special status=Illustrated;Endemic
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_110.xml
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|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_110.xml
 
|genus=Selaginella
 
|genus=Selaginella
 
|subgenus=Selaginella subg. Tetragonostachys
 
|subgenus=Selaginella subg. Tetragonostachys
 
|species=Selaginella mutica
 
|species=Selaginella mutica
|2n chromosome count=18
 
|abaxial ridge prominence=defined;defined
 
|apex architecture or shape=truncate
 
|apex shape=short-bristled;blunt
 
|base fusion=adnate
 
|base pubescence=glabrous;glabrous;pubescent
 
|base shape=decurrent;rounded
 
|branch architecture or shape=budlike
 
|bristle architecture or pubescence or relief=smooth
 
|bristle coloration=transparent;greenish transparent or whitish
 
|bristle some measurement=0.06mm;0.45mm
 
|cilium coloration=transparent
 
|cilium orientation=ascending;spreading
 
|cilium some measurement=0.03mm;0.17mm
 
|lateral branch development=determinate
 
|lateral branch shape=1-2-forked
 
|leaf architecture=monomorphic
 
|leaf coloration=green
 
|leaf fixation or orientation=appressed
 
|leaf length=1mm;2mm
 
|leaf orientation=ascending
 
|leaf shape=lanceolate;linear-lanceolate or lanceolate-elliptic
 
|leaf width=0.45mm;0.6mm
 
|main-stem development=indeterminate
 
|margin shape=ciliate;denticulate
 
|mat architecture or fragility=loose
 
|pseudowhorl arrangement=alternate
 
|pseudowhorl count=3
 
|rhizophore diameter=0.13mm;0.23mm
 
|sporophyll shape=deltate-ovate;ovate-elliptic;deltate-ovate;ovate-elliptic;ovatelanceolate
 
|stem architecture or shape=symmetric
 
|stem growth form or orientation=short-creeping
 
|stem length or size=long
 
|stem shape=forked
 
|strobilus architecture or arrangement or growth form=solitary
 
|strobilus atypical some measurement=0.6cm;1cm
 
|strobilus some measurement=1cm;3cm
 
|tip course=straight
 
 
}}<!--
 
}}<!--
  
-->[[Category:Treatment]][[Category:Selaginella subg. Tetragonostachys]]
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[[Category:Treatment]]
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[[Category:Selaginella subg. Tetragonostachys]]
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[[Category:Revised Since Print]]

Latest revision as of 17:54, 6 November 2020

Plants on rock or terrestrial, forming loose mats. Stems radially symmetric, long- to short-creeping, not readily fragmenting, ± regularly forked, without budlike arrested branches, tips straight; main stem indeterminate, lateral branches determinate, 1–2-forked. Rhizophores borne on upperside of stems, throughout stem length, 0.13–0.23 mm diam. Leaves monomorphic, in ± alternate pseudowhorls of 3, tightly appressed, ascending, green, lanceolate to linear-lanceolate or lanceolate-elliptic, 1–2 × 0.45–0.6 mm; abaxial ridges well defined; base rounded and adnate, sometimes slightly decurrent, pubescent or glabrous; margins ciliate to denticulate, cilia transparent, spreading or ascending, 0.03–0.17 mm; apex keeled, obtuse or slightly attenuate, nearly truncate in profile, blunt to short-bristled; bristle transparent to greenish transparent or whitish, smooth, 0.06–0.45 mm. Strobili solitary, (0.6–)1–3 cm; sporophylls ovate-lanceolate, ovate-elliptic, or deltate-ovate, abaxial ridges well defined, base glabrous, margins ciliate to denticulate, apex strongly to slightly keeled, short-bristled to blunt. 2n = 18.

Distribution

Ariz., Colo., N.Mex., Tex., Utah, Wyo., only in the flora.

Discussion

Selaginella mutica, S. underwoodii (R. M. Tryon 1955; C. A. Weatherby 1943), and S. wallacei all have similar patterns of variation. Study is needed to assess to what degree such variability is caused by environmental or genetic factors. Within S. mutica, two rather distinct, morphologic extremes are recognized here as varieties. Many specimens having leaves with spreading, long, marginal cilia and a short, broken, apical bristle have been considered intermediate between the two varieties, but they belong in S. mutica var. mutica.

Selaginella mutica may be one of the parent species of the putative hybrid species S. × neomexicana (see discussion). Selaginella mutica is often found growing in the same habitat with S. underwoodii, S. × neomexicana, and S. weatherbiana. According to R. M. Tryon (1955), where the two grow together, S. mutica mats gradually entirely replace mats of S. underwoodii over time. Selaginella mutica is sometimes confused with S. viridissima.

Varieties 2.

Selected References

None.

Key

1 Margins of sporophylls usually long-ciliate, seldom denticulate, cilia spreading; apex of leaves with or without bristle, 0.03–0.06 mm; leaf margins long- ciliate, rarely denticulate, cilia spreading. Selaginella mutica var. mutica
1 Margins of sporophylls mostly very short-ciliate to denticulate, cilia and teeth ascending; apex of leaves with bristle 0.2–0.45 mm; leaf margins short-ciliate to denticulate, cilia and teeth ascending. Selaginella mutica var. limitanea