Difference between revisions of "Woodsia phillipsii"

Windham

Contr. Univ. Michigan Herb. 19: 50. 1993.

Common names: Phillips's cliff fern
Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties="stem architecture or arrangement;stem orientation;stem orientation;stem orientation;stem orientation;petiole base count;petiole base duration"><b>Stems </b>compact to short-creeping, erect to horizontal, with few-to-many persistent petiole bases of unequal lengths;</span> <span class="statement" id="st-d0_s1" data-properties="scale coloration;scale coloration;scale shape;central stripe coloration;margin coloration">scales mostly uniformly brown but at least some bicolored with dark central stripe and pale-brown margins, narrowly lanceolate.</span> <span class="statement" id="st-d0_s2" data-properties="leaf length;leaf width"><b>Leaves </b>5–35 × 1.5–6 cm.</span> <span class="statement" id="st-d0_s3" data-properties="petiole coloration;petiole coloration;petiole coloration;petiole architecture;petiole fragility"><b>Petiole </b>light-brown or straw-colored when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered.</span> <span class="statement" id="st-d0_s4" data-properties="blade shape;blade architecture or shape;blade architecture or function or pubescence;blade coating"><b>Blade </b>lanceolate, usually 2-pinnate proximally, sparsely to moderately glandular, never viscid;</span> <span class="statement" id="st-d0_s5" data-properties="stalk width;tip size">glandular-hairs with thin stalks and slightly expanded tips;</span> <span class="statement" id="st-d0_s6" data-properties="glandular-hair arrangement;scale shape">rachis with scattered glandular-hairs and hairlike scales.</span> <span class="statement" id="st-d0_s7" data-properties="pinna shape;pinna shape;pinna shape;pinna length or size;pinna shape;apex shape"><b>Pinnae </b>elongate-deltate to elliptic, longer than wide, often attenuate to a narrowly acute apex;</span> <span class="statement" id="st-d0_s8" data-properties="pair count;pinnule arrangement">largest pinnae with 7–18 pairs of widely spaced pinnules;</span> <span class="statement" id="st-d0_s9" data-properties="abaxial and adaxial surface architecture or function or pubescence;hair count;hair architecture;scale count;scale architecture">abaxial and adaxial surfaces somewhat glandular, lacking nonglandular hairs or scales.</span> <span class="statement" id="st-d0_s10" data-properties="pinnule architecture or shape;pinnule shape"><b>Pinnules </b>dentate, often shallowly lobed;</span> <span class="statement" id="st-d0_s11" data-properties="margin reflectance;margin size or width;margin architecture or pubescence or shape;gland count;projection architecture;projection coloration or reflectance;tooth due-to-presence;filament architecture">margins often lustrous adaxially, somewhat thickened, with occasional glands, appearing ciliate due to presence of multicellular translucent projections on teeth that often prolonged to form twisted filaments.</span> <span class="statement" id="st-d0_s12" data-properties="vein tip size;hydathode coloration;hydathode prominence"><b>Vein </b>tips usually enlarged to form whitish hydathodes visible adaxially.</span> <span class="statement" id="st-d0_s13" data-properties="indusium length;segment size or width;segment texture;cell architecture or shape;sporangium life cycle"><b>Indusia </b>of narrow, filamentous segments, these uniseriate for most of length, composed of ± isodiametric cells, often greatly surpassing mature sporangia.</span> <span class="statement" id="st-d0_s14" data-properties="spore some measurement;2n chromosome count"><b>Spores </b>averaging 37–44 µm. <b>2n</b> = 76.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact to short-creeping, erect to horizontal, with few to many persistent petiole bases of unequal lengths; scales mostly uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. <b>Leaves</b> 5–35 × 1.5–6 cm. <b>Petiole</b> light brown or straw-colored when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered. <b>Blade</b> lanceolate, usually 2-pinnate proximally, sparsely to moderately glandular, never viscid; glandular hairs with thin stalks and slightly expanded tips; rachis with scattered glandular hairs and hairlike scales. <b>Pinnae</b> elongate-deltate to elliptic, longer than wide, often attenuate to a narrowly acute apex; largest pinnae with 7–18 pairs of widely spaced pinnules; abaxial and adaxial surfaces somewhat glandular, lacking nonglandular hairs or scales. <b>Pinnules</b> dentate, often shallowly lobed; margins often lustrous adaxially, somewhat thickened, with occasional glands, appearing ciliate due to presence of multicellular translucent projections on teeth that are often prolonged to form twisted filaments. <b>Vein</b> tips usually enlarged to form whitish hydathodes visible adaxially. <b>Indusia</b> of narrow, filamentous segments, these uniseriate for most of length, composed of ± isodiametric cells, often greatly surpassing mature sporangia. <b>Spores</b> averaging 37–44 µm. <b>2n</b> = 76.</span><!--
  
 
-->{{Treatment/Body
 
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|phenology=Sporulating summer–fall.
 
|habitat=Cliffs and rocky slopes, usually on granitic or volcanic substrates
 
|habitat=Cliffs and rocky slopes, usually on granitic or volcanic substrates
 
|elevation=1600–3200 m
 
|elevation=1600–3200 m
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|basionyms=
 
|basionyms=
 
|family=Dryopteridaceae
 
|family=Dryopteridaceae
 +
|phenology=Sporulating summer–fall.
 
|habitat=Cliffs and rocky slopes, usually on granitic or volcanic substrates
 
|habitat=Cliffs and rocky slopes, usually on granitic or volcanic substrates
 
|elevation=1600–3200 m
 
|elevation=1600–3200 m
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|publication year=1993
 
|publication year=1993
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_61.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_61.xml
 
|genus=Woodsia
 
|genus=Woodsia
 
|species=Woodsia phillipsii
 
|species=Woodsia phillipsii
|2n chromosome count=76
 
|abaxial and adaxial surface architecture or function or pubescence=glandular
 
|apex shape=acute
 
|blade architecture or function or pubescence=glandular
 
|blade architecture or shape=2-pinnate
 
|blade coating=viscid
 
|blade shape=lanceolate
 
|cell architecture or shape=isodiametric
 
|central stripe coloration=dark
 
|filament architecture=twisted
 
|gland count=occasional
 
|glandular-hair arrangement=scattered
 
|hair architecture=nonglandular
 
|hair count=lacking
 
|hydathode coloration=whitish
 
|hydathode prominence=visible
 
|indusium length=uniseriate
 
|leaf length=5cm;35cm
 
|leaf width=1.5cm;6cm
 
|margin architecture or pubescence or shape=ciliate
 
|margin coloration=pale-brown
 
|margin reflectance=lustrous
 
|margin size or width=thickened
 
|pair count=7;18
 
|petiole architecture=not articulate
 
|petiole base count=few-to-many
 
|petiole base duration=persistent
 
|petiole coloration=darker;straw-colored;light-brown
 
|petiole fragility=brittle
 
|pinna length or size=longer than wide
 
|pinna shape=attenuate;elongate-deltate;elliptic
 
|pinnule architecture or shape=dentate
 
|pinnule arrangement=spaced
 
|pinnule shape=lobed
 
|projection architecture=multicellular
 
|projection coloration or reflectance=translucent
 
|scale architecture=nonglandular
 
|scale coloration=bicolored;brown
 
|scale count=lacking
 
|scale shape=hairlike;lanceolate
 
|segment size or width=narrow
 
|segment texture=filamentous
 
|sporangium life cycle=mature
 
|spore some measurement=37um;44um
 
|stalk width=thin
 
|stem architecture or arrangement=compact
 
|stem orientation=erect;horizontal
 
|tip size=expanded
 
|tooth due-to-presence=prolonged
 
|vein tip size=enlarged
 
 
}}<!--
 
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-->[[Category:Treatment]][[Category:Woodsia]]
 
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Revision as of 14:24, 27 July 2019

Stems compact to short-creeping, erect to horizontal, with few to many persistent petiole bases of unequal lengths; scales mostly uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. Leaves 5–35 × 1.5–6 cm. Petiole light brown or straw-colored when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered. Blade lanceolate, usually 2-pinnate proximally, sparsely to moderately glandular, never viscid; glandular hairs with thin stalks and slightly expanded tips; rachis with scattered glandular hairs and hairlike scales. Pinnae elongate-deltate to elliptic, longer than wide, often attenuate to a narrowly acute apex; largest pinnae with 7–18 pairs of widely spaced pinnules; abaxial and adaxial surfaces somewhat glandular, lacking nonglandular hairs or scales. Pinnules dentate, often shallowly lobed; margins often lustrous adaxially, somewhat thickened, with occasional glands, appearing ciliate due to presence of multicellular translucent projections on teeth that are often prolonged to form twisted filaments. Vein tips usually enlarged to form whitish hydathodes visible adaxially. Indusia of narrow, filamentous segments, these uniseriate for most of length, composed of ± isodiametric cells, often greatly surpassing mature sporangia. Spores averaging 37–44 µm. 2n = 76.


Phenology: Sporulating summer–fall.
Habitat: Cliffs and rocky slopes, usually on granitic or volcanic substrates
Elevation: 1600–3200 m

Distribution

V2 61-distribution-map.gif

Ariz., N.Mex., Tex., n Mexico.

Discussion

Woodsia phillipsii traditionally has been identified as W. mexicana. It differs from typical W. mexicana, however, in having completely filamentous indusial segments, multicellular (often filamentous) translucent projections on the pinnule margins, a greater number of pinnules per pinna, and a diploid chromosome number. Woodsia phillipsii is the only diploid species currently recognized in the W. mexicana complex, and it was probably involved in the hybrid origins of both W. mexicana and W. neomexicana. Some individuals of the latter species are difficult to distinguish from W. phillipsii (see comments under W. neomexicana), and the two taxa occasionally hybridize to produce sterile triploids of intermediate morphology. Woodsia phillipsii is also known to hybridize with W. plummerae (see comments under that species) and W. cochisensis.

Selected References

None.

Lower Taxa

None.